Unranked clade
Valid Extinct

Lonchognatha

Unwin 2003

Pterosaurs are an extinct clade of flying reptiles in the order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous. Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger.

Temporal range
Triassic
Jurassic
Cretaceous
Paleogene
Neogene
252 201 145 66 0 Ma
PBDB occurrences
34
Group
Ptérosaures
Piscivore Volant Marine
Lonchognatha
click to enlarge
Mounted composite cast of Pteranodon longiceps (=P. ingens) at the American Museum of Natural History, New York. Photo credit Matt Martyniuk henteeth.com © Matt Martyniuk · CC BY 3.0 · Wikimedia
PBDB Wikipedia
Classification
Pterosauria Order
Macronychoptera Unranked clade
Novialoidea Unranked clade
Lonchognatha Unranked clade
Fossil sites 34 geolocated sites
Distribution
Top countries
🇨🇳 China
14
🇩🇪 Germany
8
🇬🇧 United Kingdom
3
🇫🇷 France
2
🇹🇿 Tanzania
2
🇦🇷 Argentina
1
🇲🇳 Mongolia
1
🇰🇿 Kazakhstan
1
🇵🇹 Portugal
1
🇺🇸 United States
1
Geological formations
Tiaojishan
12
Solnhofener Plattenkalk
5
Solnhofen
2
Cañadón Asfalto
1
Bakkar
1
Kilmaluag
1
Karabastau
1
Temporal distribution
Aptian (121.4–113.2 Ma)
1
Barremian (125.77–121.4 Ma)
1
Valanginian (137.05–132.6 Ma)
1
Tithonian (149.2–143.1 Ma)
10
Kimmeridgian (154.8–149.2 Ma)
4
Oxfordian (161.5–154.8 Ma)
12
Callovian (165.3–161.5 Ma)
1
Bathonian (168.2–165.3 Ma)
1
Aalenian (174.7–170.9 Ma)
1
Toarcian (184.2–174.7 Ma)
1
Hettangian (201.4–199.5 Ma)
1
Images 1
Bibliography
Original description
D. M. Unwin. 2003. On the phylogeny and evolutionary history of pterosaurs. Evolution and Palaeobiology of Pterosaurs, Geological Society of London, Special Publications 217:139-190 DOI ↗
Bibliography (30)
A. E. Fernandes, D. Pol, and O. W. M. Rauhut. 2024. The oldest monofenestratan pterosaur from the Queso Rallado locality (Cañadón Asfalto Formation, Toarcian) of Chubut Province, Patagonia, Argentina. Royal Society Open Science 11(12):1-20 DOI ↗
E. Martin-Silverstone, D. M. Unwin, and A. R. Cuff, E. E. Brown, L. Allington-Jones, P. M. Barrett. 2023. A new pterosaur from the Middle Jurassic of Skye, Scotland and the early diversification of flying reptiles. Journal of Vertebrate Paleontology 43(4):e2298741:1-17 DOI ↗
X.-F. Wei, R. V. Pegas, and C.-Z. Shen, Y.-F. Guo, W.-S. Ma, D.-Y. Sun, X.-Y. Zhou. 2021. Sinomacrops bondei, a new anurognathid pterosaur from the Jurassic of China and comments on the group. PeerJ 9:e11161 DOI ↗
J.-C. Lu, Q.-J. Meng, and B.-P. Wang, DLiu, C.-Z. Shen, Y.-G. Zhang. 2017. Short note on a new anurognathid pterosaur with evidence of perching behaviour from Jianchang of Liaoning Province, China. Geological Society of London Special Publications 455:95-104 DOI ↗
X.-L. Wang, S.-X. Jiang, and J.-Q. Zhang, X. Cheng, X.-F. Yu, Y.-M. Li, G.-J. Wei, X.-L. Wang. 2017. New evidence from China for the nature of the pterosaur evolutionary transition. Scientific Reports 7(42763) DOI ↗
S. Jiang, X. Wang, and X. Cheng, F. R. Costa, J. Huang, A. W. A. Kellner. 2015. Short note on an anurognathid pterosaur with a long tail from the Upper Jurassic of China. Historical Biology 27(6):718-722 DOI ↗
D. M. Martill and S. Etches. 2013. A new monofenestratan pterosaur from the Kimmeridge Clay Formation (Kimmeridgian, Upper Jurassic) of Dorset, England. Acta Palaeontologica Polonica 58(2):285-294 DOI ↗
J. Lü and D. W. E. Hone. 2012. A new Chinese anurognathid pterosaur and the evolution of pterosaurian tail lengths. Acta Palaeontologica Sinica 86(6):1317-1325 DOI ↗
J. Lü, D. M. Unwin, and D. C. Deeming, X. Jin, Y. Liu, Q. Ji. 2011. An egg-adult association, gender, and reproduction in pterosaurs. Science 331(6015):321-324 DOI ↗
C.-F. Zhou and R. R. Schoch. 2011. New material of the non-pterodactyloid pterosaur Changchengopterus pani Lü, 2009 from the Late Jurassic Tiaojishan Formation of western Liaoning. Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 260:265-275 DOI ↗
J. Lü, D. M. Unwin, and X. Jin, Y. Liu, Q. Ji. 2010. Evidence for modular evolution in a long-tailed pterosaur with a pterodactyloid skull. Proceedings of the Royal Society B 277:383-389 DOI ↗
X. Wang, A. W. A. Kellner, and S. Jiang, X. Cheng, X. Meng, T. Rodrigues. 2010. New long-tailed pterosaurs (Wukongopteridae) from western Liaoning, China. Anais da Academia Brasileira de Ciências 82(4):1045-1062 DOI ↗
J. Lü. 2009. A new non-pterodactyloid pterosaur from Qinglong County, Hebei Province of China. Acta Geologica Sinica 83(2):189-199 DOI ↗
A. Ősi and E. Prondvai. 2009. Forgotten pterosaurus in Hungarian collections: first description of Rhamphorhynchus and Pterodactylus specimens. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 252(2):167-180 DOI ↗
X. Wang, A. W. A. Kellner, and S. Jiang, X. Meng. 2009. An unusual long-tailed pterosaur with elongated neck from western Liaoning of China. Anais da Academia Brasileira de Ciências 81(4):793-812 DOI ↗
K. Carpenter, D. M. Unwin, and K. C. Cloward, C. A. Miles, C. miles. 2003. A new scaphognathine pterosaur from the Upper Jurassic Morrison Formation of Wyoming, USA. E. Buffetaut & J.-M. Mazin (eds.), Evolution and Palaeobiology of Pterosaurs, Geological Society Special Publication 217:45-54 DOI ↗
X. Wang, Z. Zhou, and F. Zhang, X. Xu. 2002. A nearly completely articulated rhamphorhynchoid pterosaur with exceptionally well-preserved wing membranes and "hairs" from Inner Mongolia, northeast China. Chinese Science Bulletin 47:226-230 DOI ↗
R. Windolf. 2000. Dinosaur sites in the Upper Jurassic of northern Germany. 5th European Workshop on Vertebrate Palaeontology. Program. Abstracts. Excursion Guides
S. -A. Ji, Q. Ji, and K. Padian. 1999. Biostratigraphy of two new pterosaurs from China. Nature 398:573-574 DOI ↗
N. N. Bakhurina and D. M. Unwin. 1995. a survey of pterosaurs from the Jurassic and Cretaceous of the former Soviet Union and Mongolia. Historical Biology 10:197-245 DOI ↗
E. Buffetaut, P. Bernier, and G. Barale, J.-P. Bourseau, C. Gaillard, J.-C. Gall, S. Wenz. 1990. A new pterosaur bone from the Kimmeridgian lithographic limestones of Cerin (France). Neues Jahrbuch für Geologie und Paläontologie, Monatshefte 1990:321-328 DOI ↗
R. A. Thulborn. 1975. Teeth of ornithischian dinosaurs from the Upper Jurassic of Portugal, with description of a hypsilophodontid (Phyllodon henkeli gen. et sp. nov.) from the Guimarota lignite. Contribuição para o conhecimento da Fauna do Kimerridgiano da Mina de Lignito Guimarota (Leiria, Portugal). Serviços Geológicos de Portugal, Memória (Nova Série) 22:89-134
P. Wellnhofer. 1975. Die Rhamphorhynchoidea (Pterosauria) der Oberjura-Plattenkalke Süddeutschlands, Teil II, Systematische Beschreibung. Palaeontographica Abteilung A 148:132-186
P. Wellnhofer. 1970. Die Pterodactyloidea (Pterosauria) der Oberjura-Plattenkalke Süddeutschlands [The Pterydactyloidea (Pterosauria) of the Upper Jurassic plattenkalks of southern Germany]. Bayerische Akademie der Wissenschaften Mathematisch-Naturwissenschaftliche Klasse Abhandlungen, Neue Folge 141:1-133
A. N. Riabinin. 1948. Zametka o letayushchem yashchere iz yury Kara-Tau [Note about a flying reptile from the Jurassic of Kara-Tau]. Trudy Paleontologicheskogo Instituta Akademiy Nauk SSSR 1:86-93
H. Reck. 1931. Die deutschostafrikanischen Flugsaurier [The German East African pterosaurs]. Centralblatt für Mineralogie, Geologie und Paläontologie, Abteilung B: Geologie und Paläontologie 7:321-336
R. Lydekker. 1888. Catalogue of the Fossil Reptilia and Amphibia in the British Museum (Natural History). Part I. Containing the Orders Ornithosauria, Crocodilia, Dinosauria, Squamata, Rhynchocephalia, and Proterosauria. British Museum (Natural History), London DOI ↗
R. Etheridge. 1885. Manual of Geology: Theoretical and Practical. Part II. Stratigraphical Geology and Palaeontology. Charles Griffin and Company, London DOI ↗
P. Gervais. 1852. Zoologie et Paléontologie Françaises (Animaux Vertébrés) ou Nouvelles Recherches sur les Animaux Vivants et Fossiles de la France. 1–3:1-274+ DOI ↗
S. T. Soemmerring. 1812. Über einen Ornithocephalus [On an Ornithocephalus]. Denkschriften der Königlichen Akademie der Wissenschaften zu München 3:89-158