Maniraptora

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Diuqin is an unenlagiine dinosaur from the Late Cretaceous of what is now Argentina. Unenlagiines, known exclusively from South America, are usually classified as dromaeosaurs though this is sometimes debated. Like dromaeosaurs, they were covered in feathers, carnivorous, and had the large sickle-like claw on the inner toe of each foot. Unique to unenlagiines is their elongated snout, suggesting a piscivorous diet. Diuqin was a medium-sized unenlagiine, at about 4 m in length.

Diuqin is an unenlagiine dinosaur from the Late Cretaceous of what is now Argentina. Unenlagiines, known exclusively from South America, are usually classified as dromaeosaurs though this is sometimes debated. Like dromaeosaurs, they were covered in feathers, carnivorous, and had the large sickle-like claw on the inner toe of each foot. Unique to unenlagiines is their elongated snout, suggesting a piscivorous diet. Diuqin was a medium-sized unenlagiine, at about 4 m in length.

claw feather Argentina Cretaceous +5
Figure 1. Evolution of macroecological traits in Dinosauria. Large scale event in dinosaur evolution (a); the origin of dinosaurs (star), hyperthermals (volcano), the earliest fossil Avialae (bird), the earliest fossil angiosperm (flower), the Cretaceous/Palaeogene mass extinction (asteroid). Phylogeny of dinosaurs (b) redrawn from Sereno and adapted to the current consensus and upon which an ancestral state reconstruction of temperature niche (mean annual temperature) after Chiarenza et al. is plotted; Mesozoic palaeogeographies (c) for Triassic (T), Jurassic (J) and Cretaceous (K). Silhouette colours symbolize body mass for each of the taxa represented; information on dietary habits are plotted after Barrett and Zanno & Makovicky; numbers represent clades discussed through this study: 1, Ornithischia; 2, Thyreophora; 3, Ornithopoda; 4, Hadrosauroidea; 5, Marginocephalia; 6, Ceratopsia; 7, Saurischia; 8, Sauropodomorpha; 9, Sauropoda; 10, Theropoda; 11, Ceratosauria; 12, Tetanurae; 13, Coelurosauria; 14, Maniraptoriformes; 15, Maniraptora; 16, Deinonychosauria; 17, Avialae; 18, Ornithothoraces. Palaeogeographies modified from original plots via R package ‘mapast’ using plate models by Scotese.

Figure 1. Evolution of macroecological traits in Dinosauria. Large scale event in dinosaur evolution (a); the origin of dinosaurs (star), hyperthermals (volcano), the earliest fossil Avialae (bird), the earliest fossil angiosperm (flower), the Cretaceous/Palaeogene mass extinction (asteroid). Phylogeny of dinosaurs (b) redrawn from Sereno and adapted to the current consensus and upon which an ancestral state reconstruction of temperature niche (mean annual temperature) after Chiarenza et al. is plotted; Mesozoic palaeogeographies (c) for Triassic (T), Jurassic (J) and Cretaceous (K). Silhouette colours symbolize body mass for each of the taxa represented; information on dietary habits are plotted after Barrett and Zanno & Makovicky; numbers represent clades discussed through this study: 1, Ornithischia; 2, Thyreophora; 3, Ornithopoda; 4, Hadrosauroidea; 5, Marginocephalia; 6, Ceratopsia; 7, Saurischia; 8, Sauropodomorpha; 9, Sauropoda; 10, Theropoda; 11, Ceratosauria; 12, Tetanurae; 13, Coelurosauria; 14, Maniraptoriformes; 15, Maniraptora; 16, Deinonychosauria; 17, Avialae; 18, Ornithothoraces. Palaeogeographies modified from original plots via R package ‘mapast’ using plate models by Scotese.

scale Cretaceous Jurassic Mesozoic +23
Representative taxa from the Santonian Iharkút fauna from the Csehbánya Formation, Bakony Mountains, western Hungary. A Pannoniasaurus inexpectatus (Squamata, Mosasauroidea), dorsal vertebra (MTM uncatalogued) in dorsal view (photo by Réka Kalmár) B Foxemys trabanti (Pleurodira, Bothremydidae), skull (MTM V 2010.215.1.) in dorsal view (photo by Márton Rabi). C Bicuspidon aff. hatzegiensis (Squamata, Borioteiioidea), left dentary (MTM 2006.112.1.) in medial view (photo by László Makádi) D Basal tetanuran (Theropoda, Tetanurae), tooth (MTM V.01.54) in ?lingual view E Indeterminate abelisaurid (Theropoda, Abelisauridae), pedal ungual phalanx (MTM V 2008.43.1.) in lateral view F Pneumatoraptor fodori (Theropoda, Paraves), left scapulocoracoid (holotype, MTM V 2008.38.1.) in lateral view G Mochlodon vorosi (Ornithopoda, Rhabdodontidae), left dentary (holotype, MTM V 2010.105.1) in lateral view H Bakonydraco galaczi (Pterosauria, Azhdarchidae), mandible (holotype, MTM 2007.110.1) in dorsal view I Iharkutosuchus makadii (Eusuchia, Hylaeochampsidae), skull (holotype, MTM 2006.52.1) in dorsal view J Hungarosaurus tormai (Ankylosauria, Nodosauridae), right dentary (MTM 2007.25.2) in lateral view K Bauxitornis mindszentyae (Aves, Enantiornithes), left tarsometatarsus (holotype, MTM V 2009.38.1) in anterior view L Ajkaceratops kozmai (Ceratopsia), fused rostral and premaxillae (holotype, MTM V 2009.192.1) in lateral view. Scale bars: 2 cm in A, V, G, H, I, J; 1 cm in D, E, F, K, L; 1 mm in C.

Representative taxa from the Santonian Iharkút fauna from the Csehbánya Formation, Bakony Mountains, western Hungary. A Pannoniasaurus inexpectatus (Squamata, Mosasauroidea), dorsal vertebra (MTM uncatalogued) in dorsal view (photo by Réka Kalmár) B Foxemys trabanti (Pleurodira, Bothremydidae), skull (MTM V 2010.215.1.) in dorsal view (photo by Márton Rabi). C Bicuspidon aff. hatzegiensis (Squamata, Borioteiioidea), left dentary (MTM 2006.112.1.) in medial view (photo by László Makádi) D Basal tetanuran (Theropoda, Tetanurae), tooth (MTM V.01.54) in ?lingual view E Indeterminate abelisaurid (Theropoda, Abelisauridae), pedal ungual phalanx (MTM V 2008.43.1.) in lateral view F Pneumatoraptor fodori (Theropoda, Paraves), left scapulocoracoid (holotype, MTM V 2008.38.1.) in lateral view G Mochlodon vorosi (Ornithopoda, Rhabdodontidae), left dentary (holotype, MTM V 2010.105.1) in lateral view H Bakonydraco galaczi (Pterosauria, Azhdarchidae), mandible (holotype, MTM 2007.110.1) in dorsal view I Iharkutosuchus makadii (Eusuchia, Hylaeochampsidae), skull (holotype, MTM 2006.52.1) in dorsal view J Hungarosaurus tormai (Ankylosauria, Nodosauridae), right dentary (MTM 2007.25.2) in lateral view K Bauxitornis mindszentyae (Aves, Enantiornithes), left tarsometatarsus (holotype, MTM V 2009.38.1) in anterior view L Ajkaceratops kozmai (Ceratopsia), fused rostral and premaxillae (holotype, MTM V 2009.192.1) in lateral view. Scale bars: 2 cm in A, V, G, H, I, J; 1 cm in D, E, F, K, L; 1 mm in C.

tooth vertebra Hungary Santonian +15
Life restoration of Chiappeavis magnapremaxillo.

Life restoration of Chiappeavis magnapremaxillo.

Maniraptora
Archaeopteryx, Romanian stamp from 1993

Archaeopteryx, Romanian stamp from 1993

Archaeopteryx Maniraptora
Life reconstruction of Alexornis antecedens.

Life reconstruction of Alexornis antecedens.

Maniraptora
Composite figure of various feathered maniraptorans: Archaeopteryx, Anchiornis, Microraptor and Zhenyuanlong

Composite figure of various feathered maniraptorans: Archaeopteryx, Anchiornis, Microraptor and Zhenyuanlong

Anchiornis Archaeopteryx Coelurosauria Maniraptora +2
Cladogram of amniotes. Based on File:Cladogram Amniota A.jpg

Cladogram of amniotes. Based on File:Cladogram Amniota A.jpg

Coelurosauria Dinosauria Maniraptora Ornithischia +2
Cladogram of the former "Reptilia" (red)

Cladogram of the former "Reptilia" (red)

Coelurosauria Dinosauria Maniraptora Ornithischia +2
Six maniraptoran dinosaurs (top to bottom):  the Zamyn Khondt oviraptorosaur, Sinornithosaurus, the Alaskan troodontid, Patagonykus, a short-tailed albatross, Nothronychus
Taxa Maniraptora

Six maniraptoran dinosaurs (top to bottom): the Zamyn Khondt oviraptorosaur, Sinornithosaurus, the Alaskan troodontid, Patagonykus, a short-tailed albatross, Nothronychus

Coelurosauria Dinosauria Maniraptora Nothronychus +4
Six exemplar coelurosaurs (clockwise from top left): Tyrannosaurus rex, Sinosauropteryx prima, Deinonychus antirrhopus, Archaeopteryx lithographica, an undescribed crested oviraptorid, and Passer domesticus.  This is a collection of six different works which have been previously published on Wikimedia Commons (see source field below).
Taxa Coelurosauria

Six exemplar coelurosaurs (clockwise from top left): Tyrannosaurus rex, Sinosauropteryx prima, Deinonychus antirrhopus, Archaeopteryx lithographica, an undescribed crested oviraptorid, and Passer domesticus. This is a collection of six different works which have been previously published on Wikimedia Commons (see source field below).

Archaeopteryx Bullatosauria Coelurosauria Deinocheirosauria +9
Six exemplar coelurosaurs (clockwise from top left): Tyrannosaurus rex, Sinosauropteryx prima, Deinonychus antirrhopus, Archaeopteryx lithographica, an undescribed crested oviraptorid, and Passer domesticus.  This is a collection of six different works which have been previously published on Wikimedia Commons (see source field below).
Taxa Maniraptoriformes

Six exemplar coelurosaurs (clockwise from top left): Tyrannosaurus rex, Sinosauropteryx prima, Deinonychus antirrhopus, Archaeopteryx lithographica, an undescribed crested oviraptorid, and Passer domesticus. This is a collection of six different works which have been previously published on Wikimedia Commons (see source field below).

Archaeopteryx Bullatosauria Coelurosauria Deinocheirosauria +9
Six exemplar coelurosaurs (clockwise from top left): Tyrannosaurus rex, Sinosauropteryx prima, Deinonychus antirrhopus, Archaeopteryx lithographica, an undescribed crested oviraptorid, and Passer domesticus.  This is a collection of six different works which have been previously published on Wikimedia Commons (see source field below).
Taxa Maniraptoromorpha

Six exemplar coelurosaurs (clockwise from top left): Tyrannosaurus rex, Sinosauropteryx prima, Deinonychus antirrhopus, Archaeopteryx lithographica, an undescribed crested oviraptorid, and Passer domesticus. This is a collection of six different works which have been previously published on Wikimedia Commons (see source field below).

Archaeopteryx Bullatosauria Coelurosauria Deinocheirosauria +9
Six exemplar coelurosaurs (clockwise from top left): Tyrannosaurus rex, Sinosauropteryx prima, Deinonychus antirrhopus, Archaeopteryx lithographica, an undescribed crested oviraptorid, and Passer domesticus.  This is a collection of six different works which have been previously published on Wikimedia Commons (see source field below).
Taxa Tyrannoraptora

Six exemplar coelurosaurs (clockwise from top left): Tyrannosaurus rex, Sinosauropteryx prima, Deinonychus antirrhopus, Archaeopteryx lithographica, an undescribed crested oviraptorid, and Passer domesticus. This is a collection of six different works which have been previously published on Wikimedia Commons (see source field below).

Archaeopteryx Bullatosauria Coelurosauria Deinocheirosauria +9
Six exemplar coelurosaurs (clockwise from top left): Tyrannosaurus rex, Sinosauropteryx prima, Deinonychus antirrhopus, Archaeopteryx lithographica, an undescribed crested oviraptorid, and Passer domesticus.  This is a collection of six different works which have been previously published on Wikimedia Commons (see source field below).
Taxa Bullatosauria

Six exemplar coelurosaurs (clockwise from top left): Tyrannosaurus rex, Sinosauropteryx prima, Deinonychus antirrhopus, Archaeopteryx lithographica, an undescribed crested oviraptorid, and Passer domesticus. This is a collection of six different works which have been previously published on Wikimedia Commons (see source field below).

Archaeopteryx Bullatosauria Coelurosauria Deinocheirosauria +9
Six exemplar coelurosaurs (clockwise from top left): Tyrannosaurus rex, Sinosauropteryx prima, Deinonychus antirrhopus, Archaeopteryx lithographica, an undescribed crested oviraptorid, and Passer domesticus.  This is a collection of six different works which have been previously published on Wikimedia Commons (see source field below).
Taxa Protoavia

Six exemplar coelurosaurs (clockwise from top left): Tyrannosaurus rex, Sinosauropteryx prima, Deinonychus antirrhopus, Archaeopteryx lithographica, an undescribed crested oviraptorid, and Passer domesticus. This is a collection of six different works which have been previously published on Wikimedia Commons (see source field below).

Archaeopteryx Bullatosauria Coelurosauria Deinocheirosauria +9