Ichthyosauria

Taxon

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Various fossils pertaining to the holotype of the Triassic ichthyosaur Toretocnemus (originally Leptocheirus, also Merriamia) zitteli. This image is derived from plate 23 in Merriam (1903), done by an uncredited artist. The arrangement of the individual figures has been modified from the original (most notably, figures 3 and 4 were swapped to minimize confusion).
Original description:
Leptocheirus zitteli n. gen. and sp.
Figures reproduced natural size from the type specimen.

Fig. 1.— Right side of skull.
Fig. 2. — Cross-section of posterior portion of the lower jaw, taken above the point marked A on the lateral view of the skull.
Fig. 3. — Cross-section of the upper and lower jaws, taken at the break immediately behind the point marked Sp on the lateral view of the skull.
Fig. 4. — Lateral view of an anterior caudal centrum.
Legend (modified from original):

Ar - articular
A - angular
D - dentary
J - jugal
L - lacrimal
Mx - maxilla
Po - postorbital
Sa - surangular
Se - sclerotic ring
Sp - spenial
T - cross-section of tooth
X - doubtful element

Various fossils pertaining to the holotype of the Triassic ichthyosaur Toretocnemus (originally Leptocheirus, also Merriamia) zitteli. This image is derived from plate 23 in Merriam (1903), done by an uncredited artist. The arrangement of the individual figures has been modified from the original (most notably, figures 3 and 4 were swapped to minimize confusion). Original description: Leptocheirus zitteli n. gen. and sp. Figures reproduced natural size from the type specimen. Fig. 1.— Right side of skull. Fig. 2. — Cross-section of posterior portion of the lower jaw, taken above the point marked A on the lateral view of the skull. Fig. 3. — Cross-section of the upper and lower jaws, taken at the break immediately behind the point marked Sp on the lateral view of the skull. Fig. 4. — Lateral view of an anterior caudal centrum. Legend (modified from original): Ar - articular A - angular D - dentary J - jugal L - lacrimal Mx - maxilla Po - postorbital Sa - surangular Se - sclerotic ring Sp - spenial T - cross-section of tooth X - doubtful element

description Trias fossile holotype +7
The shoulder girdle of the holotype of the Triassic ichthyosaur Toretocnemus (originally Leptocheirus, also Merriamia) zitteli. This image is derived from plate 21 in Merriam (1903), done by an uncredited artist. The arrangement of the individual figures has been slightly modified from the original.
Original description: Leptocheirus zitteli n. gen. and sp.
Figures reproduced natural size from the type specimen.

Fig. 1.—Clavicles (cl) and probable interelavicle (Ic and Ic')- The interclavicle has been separated into two fragments by a crack in the matrix.
Fig. 2.—Inner side of the coracoids and the left scapula.

The shoulder girdle of the holotype of the Triassic ichthyosaur Toretocnemus (originally Leptocheirus, also Merriamia) zitteli. This image is derived from plate 21 in Merriam (1903), done by an uncredited artist. The arrangement of the individual figures has been slightly modified from the original. Original description: Leptocheirus zitteli n. gen. and sp. Figures reproduced natural size from the type specimen. Fig. 1.—Clavicles (cl) and probable interelavicle (Ic and Ic')- The interclavicle has been separated into two fragments by a crack in the matrix. Fig. 2.—Inner side of the coracoids and the left scapula.

description Trias holotype spécimen +5
Forelimb and partial hindlimb of the holotype of the Triassic ichthyosaur Toretocnemus (originally Leptocheirus, also Merriamia) zitteli. This image is derived from plate 22 in Merriam (1903), done by an uncredited artist.
Original description: Leptocheirus zitteli n. gen. and sp.
Figures reproduced natural size from the type specimen.

Fig. 1. — Superior side of left anterior limb. r, radius.
Fig. 2. — Posterior limb. t, tibia.

Forelimb and partial hindlimb of the holotype of the Triassic ichthyosaur Toretocnemus (originally Leptocheirus, also Merriamia) zitteli. This image is derived from plate 22 in Merriam (1903), done by an uncredited artist. Original description: Leptocheirus zitteli n. gen. and sp. Figures reproduced natural size from the type specimen. Fig. 1. — Superior side of left anterior limb. r, radius. Fig. 2. — Posterior limb. t, tibia.

membre description Trias holotype +7
Various fossils pertaining to the holotype of the Triassic ichthyosaur Toretocnemus californicus. This image is derived from plate 24 in Merriam (1903), done by an uncredited artist. The arrangement of the individual figures has been modified from the original.
Original description:
Toretocnemus californicus n. gen. and sp.
Figures reproduced natural size from the type specimen.

Fig. 1. — Inferior side of right posterior limb. t, tibia.
Fig. 2. — Right anterior limb. r, radius.
Fig. 3.— Pelvic arch.
Fig. 4. — Middle dorsal vertebrae and a rib from the same region.

Various fossils pertaining to the holotype of the Triassic ichthyosaur Toretocnemus californicus. This image is derived from plate 24 in Merriam (1903), done by an uncredited artist. The arrangement of the individual figures has been modified from the original. Original description: Toretocnemus californicus n. gen. and sp. Figures reproduced natural size from the type specimen. Fig. 1. — Inferior side of right posterior limb. t, tibia. Fig. 2. — Right anterior limb. r, radius. Fig. 3.— Pelvic arch. Fig. 4. — Middle dorsal vertebrae and a rib from the same region.

membre description Trias fossile +6
Illustration of some anterior caudal vertebrae of the Californian ichthyosaur Torteocnemus californicus (specimen UCMP 8100), from Merriam's 1908 monograph on American Triassic ichthyosaurs. This illustration appeared as figure 4 of plate 14; it has been modified to remove the stamp placed on top of it. While Merriam credits two illustrators as having produced artwork for this publication, it is not clear which of them created this image.

Illustration of some anterior caudal vertebrae of the Californian ichthyosaur Torteocnemus californicus (specimen UCMP 8100), from Merriam's 1908 monograph on American Triassic ichthyosaurs. This illustration appeared as figure 4 of plate 14; it has been modified to remove the stamp placed on top of it. While Merriam credits two illustrators as having produced artwork for this publication, it is not clear which of them created this image.

Trias spécimen Ichthyosauria Merriamia +1
Life restoration of the Triassic ichthyosaur Qianichthyosaurus xingyiensis.
References
Motani, Ryosuke; Jiang, Da-yong; Tintori, Andrea; Ji, Cheng; Huang, Jian-dong (2017). "Pre- versus post-mass extinction divergence of Mesozoic marine reptiles dictated by time-scale dependence of evolutionary rates". Proceedings of the Royal Society B 284 (1854): 20170241. DOI:10.1098/rspb.2017.0241.

Life restoration of the Triassic ichthyosaur Qianichthyosaurus xingyiensis. References Motani, Ryosuke; Jiang, Da-yong; Tintori, Andrea; Ji, Cheng; Huang, Jian-dong (2017). "Pre- versus post-mass extinction divergence of Mesozoic marine reptiles dictated by time-scale dependence of evolutionary rates". Proceedings of the Royal Society B 284 (1854): 20170241. DOI:10.1098/rspb.2017.0241.

écaille Mésozoïque Trias Ichthyosauria +3
Comparison between the tails of for different Ichthyosauria specimens, captions present and in German, from Abel, O. (1919). Die Stämme der Wirbeltiere.

Comparison between the tails of for different Ichthyosauria specimens, captions present and in German, from Abel, O. (1919). Die Stämme der Wirbeltiere.

spécimen Ichthyosauria Mixosauridae
Figure description from paper: "Figure 23: Early and Middle Triassic ichthyopterygian heads possessing longirostry.
Simplified outlines of four different Early and Middle Triassic ichthyopterygian heads possessing a long and slender rostrum. Specimens are at the same scale. (A) Utatsusaurus hataii (UHR 30691, Motani, Minoura & Ando, 1998); (B) Grippia longirostris (PMU R445, Motani, 2000); (C) Mixosaurus cornalianus (BES SC 1000, Renesto et al., 2020); (D) Besanosaurus leptorhynchus (PIMUZ T 4847, this paper); (E) Cymbospondylus buchseri (PIMUZ T 4351, Sander, 1989). Scale bar represents 10 cm."
References:

Bindellini, G.; Wolniewicz, A.S.; Miedema, F.; Scheyer, T.M.; Dal Sasso, C. (2021). "Cranial anatomy of Besanosaurus leptorhynchus Dal Sasso & Pinna, 1996 (Reptilia: Ichthyosauria) from the Middle Triassic Besano Formation of Monte San Giorgio, Italy/Switzerland: Taxonomic and palaeobiological implications". PeerJ 9: e11179. DOI:10.7717/peerj.11179.

Figure description from paper: "Figure 23: Early and Middle Triassic ichthyopterygian heads possessing longirostry. Simplified outlines of four different Early and Middle Triassic ichthyopterygian heads possessing a long and slender rostrum. Specimens are at the same scale. (A) Utatsusaurus hataii (UHR 30691, Motani, Minoura & Ando, 1998); (B) Grippia longirostris (PMU R445, Motani, 2000); (C) Mixosaurus cornalianus (BES SC 1000, Renesto et al., 2020); (D) Besanosaurus leptorhynchus (PIMUZ T 4847, this paper); (E) Cymbospondylus buchseri (PIMUZ T 4351, Sander, 1989). Scale bar represents 10 cm." References: Bindellini, G.; Wolniewicz, A.S.; Miedema, F.; Scheyer, T.M.; Dal Sasso, C. (2021). "Cranial anatomy of Besanosaurus leptorhynchus Dal Sasso & Pinna, 1996 (Reptilia: Ichthyosauria) from the Middle Triassic Besano Formation of Monte San Giorgio, Italy/Switzerland: Taxonomic and palaeobiological implications". PeerJ 9: e11179. DOI:10.7717/peerj.11179.

écaille description Italie Suisse +12
Besanosaurus leptorhynchus referred specimen PIMUZ T 4376, preserved on the same slab as a specimen of the smaller Mixosaurus. Scale bar represents 50 cm (1.6 ft). Figure 3C of Bindellini et al. (2021).
References:

Bindellini, G.; Wolniewicz, A.S.; Miedema, F.; Scheyer, T.M.; Dal Sasso, C. (2021). "Cranial anatomy of Besanosaurus leptorhynchus Dal Sasso & Pinna, 1996 (Reptilia: Ichthyosauria) from the Middle Triassic Besano Formation of Monte San Giorgio, Italy/Switzerland: Taxonomic and palaeobiological implications". PeerJ 9: e11179. DOI:10.7717/peerj.11179.

Besanosaurus leptorhynchus referred specimen PIMUZ T 4376, preserved on the same slab as a specimen of the smaller Mixosaurus. Scale bar represents 50 cm (1.6 ft). Figure 3C of Bindellini et al. (2021). References: Bindellini, G.; Wolniewicz, A.S.; Miedema, F.; Scheyer, T.M.; Dal Sasso, C. (2021). "Cranial anatomy of Besanosaurus leptorhynchus Dal Sasso & Pinna, 1996 (Reptilia: Ichthyosauria) from the Middle Triassic Besano Formation of Monte San Giorgio, Italy/Switzerland: Taxonomic and palaeobiological implications". PeerJ 9: e11179. DOI:10.7717/peerj.11179.

écaille Italie Suisse Trias moyen +8
Modified version of figure 23 of Bindellini et al. (2021) showing just the ichthyosaurs from the Besano Formation: Besanosaurus leptorhynchus (A), Mixosaurus cornalianus (B), and Cymbospondylus buchseri (C). The original figure was uploaded unmodified here: File:Early to Middle Triassic Longirostrine Ichthyopterygians.png
References:

Bindellini, G.; Wolniewicz, A.S.; Miedema, F.; Scheyer, T.M.; Dal Sasso, C. (2021). "Cranial anatomy of Besanosaurus leptorhynchus Dal Sasso & Pinna, 1996 (Reptilia: Ichthyosauria) from the Middle Triassic Besano Formation of Monte San Giorgio, Italy/Switzerland: Taxonomic and palaeobiological implications". PeerJ 9: e11179. DOI:10.7717/peerj.11179.

Modified version of figure 23 of Bindellini et al. (2021) showing just the ichthyosaurs from the Besano Formation: Besanosaurus leptorhynchus (A), Mixosaurus cornalianus (B), and Cymbospondylus buchseri (C). The original figure was uploaded unmodified here: File:Early to Middle Triassic Longirostrine Ichthyopterygians.png References: Bindellini, G.; Wolniewicz, A.S.; Miedema, F.; Scheyer, T.M.; Dal Sasso, C. (2021). "Cranial anatomy of Besanosaurus leptorhynchus Dal Sasso & Pinna, 1996 (Reptilia: Ichthyosauria) from the Middle Triassic Besano Formation of Monte San Giorgio, Italy/Switzerland: Taxonomic and palaeobiological implications". PeerJ 9: e11179. DOI:10.7717/peerj.11179.

Italie Suisse Trias moyen Trias +7
Identifier: waterreptilesofp1914will
Title: Water reptiles of the past and present
Year: 1914 (1910s)
Authors:  Williston, Samuel Wendell, 1851-1918
Subjects:  Aquatic reptiles
Publisher:  Chicago, Ill., The University of Chicago Press
Contributing Library:  Boston Public Library
Digitizing Sponsor:  Boston Public Library

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Text Appearing Before Image:
served and very complete skeletons ofdifferent species of ichthyosaurs from the Jurassic deposits ofWiirtemberg, in which remains of these animals occur in great 112 WATER REPTILES OF THE PAST AND PRESENT profusion. His researches, and those of several authors since then,supplementing and confirming or disproving those of the manyobservers made during the preceding seventy years, have finallydetermined almost perfectly the complete structure of the moretypical ichthyosaurs, enabling us to infer not a little as to theirhabits and distribution in the old Jurassic oceans. Within thepast few years the discoveries of Professor J. C. Merriam of Cali-fornia have likewise added greatly to our knowledge of the earlierichthyosaurs. It may now truthfully be said that of no group ofextinct reptiles do we have a more complete and satisfactory knowl-edge than of the ichthyosaurs. Nevertheless we have yet very much more to learn about theorder Ichthyosauria as a whole—whence they came and how they
Text Appearing After Image:
Fig. 52.—Ichthyosaurus quadricissus.museum, from Dr. Dreverman. Photograph of specimen in Senckenberg originated; what their nearest kin were among other reptiles; andespecially, more about the connecting links between them andterrestrial reptiles. They have, as an order, so isolated a position,are so widely separated from all other reptiles in structure, that theyhave long been a puzzle to paleontologists. Like the whales andother cetaceans among mammals, we know the ichthyosaurs wellin the plenitude of their power and the fulness of their development,but have yet only an imperfect knowledge of their earlier history,and none whatever of their earliest. However, as will be seenfarther on, the recent discoveries by Merriam have shed much lighton some of the stages of their evolution. So nearly perfectly wereall the later ichthyosaurs adapted to their life in the water that itwas believed by nearly all paleontologists until about a score of years ICHTHYOSAURIA 3 ago that they had desc

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Please note that these images are extracted from scanned page images that may have been digitally enhanced for readability - coloration and appearance of these illustrations may not perfectly resemble the original work.

Identifier: waterreptilesofp1914will Title: Water reptiles of the past and present Year: 1914 (1910s) Authors: Williston, Samuel Wendell, 1851-1918 Subjects: Aquatic reptiles Publisher: Chicago, Ill., The University of Chicago Press Contributing Library: Boston Public Library Digitizing Sponsor: Boston Public Library View Book Page: Book Viewer About This Book: Catalog Entry View All Images: All Images From Book Click here to view book online to see this illustration in context in a browseable online version of this book. Text Appearing Before Image: served and very complete skeletons ofdifferent species of ichthyosaurs from the Jurassic deposits ofWiirtemberg, in which remains of these animals occur in great 112 WATER REPTILES OF THE PAST AND PRESENT profusion. His researches, and those of several authors since then,supplementing and confirming or disproving those of the manyobservers made during the preceding seventy years, have finallydetermined almost perfectly the complete structure of the moretypical ichthyosaurs, enabling us to infer not a little as to theirhabits and distribution in the old Jurassic oceans. Within thepast few years the discoveries of Professor J. C. Merriam of Cali-fornia have likewise added greatly to our knowledge of the earlierichthyosaurs. It may now truthfully be said that of no group ofextinct reptiles do we have a more complete and satisfactory knowl-edge than of the ichthyosaurs. Nevertheless we have yet very much more to learn about theorder Ichthyosauria as a whole—whence they came and how they Text Appearing After Image: Fig. 52.—Ichthyosaurus quadricissus.museum, from Dr. Dreverman. Photograph of specimen in Senckenberg originated; what their nearest kin were among other reptiles; andespecially, more about the connecting links between them andterrestrial reptiles. They have, as an order, so isolated a position,are so widely separated from all other reptiles in structure, that theyhave long been a puzzle to paleontologists. Like the whales andother cetaceans among mammals, we know the ichthyosaurs wellin the plenitude of their power and the fulness of their development,but have yet only an imperfect knowledge of their earlier history,and none whatever of their earliest. However, as will be seenfarther on, the recent discoveries by Merriam have shed much lighton some of the stages of their evolution. So nearly perfectly wereall the later ichthyosaurs adapted to their life in the water that itwas believed by nearly all paleontologists until about a score of years ICHTHYOSAURIA 3 ago that they had desc Note About Images Please note that these images are extracted from scanned page images that may have been digitally enhanced for readability - coloration and appearance of these illustrations may not perfectly resemble the original work.

Jurassique Ichthyosauria Stenopterygiidae squelette
Utatsusaurus hataii, an Early Triassic Ichthyosaur from Japan, pencil drawing
Taxons Grippiidae

Utatsusaurus hataii, an Early Triassic Ichthyosaur from Japan, pencil drawing

dessin Japon Early Triassic Trias +4
Skull of the Middle Triassic North-American ichthyosaur Cymbospondylus petrinus, preserved in three dimensions. This specimen is cataloged under the number UCMP 9913. More information is available from the source link.
Taxons Longipinnati

Skull of the Middle Triassic North-American ichthyosaur Cymbospondylus petrinus, preserved in three dimensions. This specimen is cataloged under the number UCMP 9913. More information is available from the source link.

musée Trias moyen Trias spécimen +5
Ichthyosauria
Bituminous claystone, Lower Jurassic, Hesselberg, Middle Franconia.
Sedimentary rock of very thin laminated layers (particles less than 0,0002 mm).

Rock formation is known for its rich fossil findings, ex. Ichthyosaur.

Bituminous claystone, Lower Jurassic, Hesselberg, Middle Franconia. Sedimentary rock of very thin laminated layers (particles less than 0,0002 mm). Rock formation is known for its rich fossil findings, ex. Ichthyosaur.

Jurassique fossile Ichthyosauria formation
Life restoration of the German Jurassic ichthyosaur Suevoleviathan disinteger. The dorsal and caudal fins are loosely based on those of Stenopterygius.
References
Maisch, M.W. (2020). "The best-preserved skeleton of Suevoleviathan integer (Bronn, 1844)(Reptilia: Ichthyosauria) from the lower Jurassic of south-western Germany, with a discussion of the genus". Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 297 (2): 153–172.
Maisch, M.W. (1998). "A new ichthyosaur genus from the Posidonia Shale (Lower Toarcian, Jurassic) of Holzmaden, SW-Germany with comments on the phylogeny of post-Triassic ichthyosaurs". Neues Jahrbuch für Geologie und Paläontologie-Abhandlungen 209 (1): 47–78.
Taxons Suevoleviathan

Life restoration of the German Jurassic ichthyosaur Suevoleviathan disinteger. The dorsal and caudal fins are loosely based on those of Stenopterygius. References Maisch, M.W. (2020). "The best-preserved skeleton of Suevoleviathan integer (Bronn, 1844)(Reptilia: Ichthyosauria) from the lower Jurassic of south-western Germany, with a discussion of the genus". Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 297 (2): 153–172. Maisch, M.W. (1998). "A new ichthyosaur genus from the Posidonia Shale (Lower Toarcian, Jurassic) of Holzmaden, SW-Germany with comments on the phylogeny of post-Triassic ichthyosaurs". Neues Jahrbuch für Geologie und Paläontologie-Abhandlungen 209 (1): 47–78.

Allemagne Posidonia Shale Jurassique Toarcien +6
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Actualités

On le croyait roi des océans au Jurassique : une vertèbre percée raconte un combat d’une violence extrême
On le croyait roi des océans au Jurassique : une vertèbre percée raconte un combat d’une violence extrême
os dent vertèbre chasse musée Jurassique fossile Ichthyosauria Ichthyosaurus Pliosaurus comportement
Oublié pendant plus d’un siècle dans les tiroirs d’un musée, un fossile refait aujourd’hui surface et raconte une scène de violence extrême dans les océans du Jurassique. Une vertèbre d’ichtyosaure, marquée par l’impact d’une dent brisée, révèle peut-être l’attaque d’un des plus redoutables...
17/06/2026 futura-terre
Un gigantesque « dragon des mers » vieux de 180 millions d’années découvert au Royaume-Uni
Un gigantesque « dragon des mers » vieux de 180 millions d’années découvert au Royaume-Uni
Royaume-Uni fossile Ichthyosauria découverte
180 millions d'années sous nos pieds. Un fossile d'ichtyosaure colossal vient d'être mis au jour en Angleterre, et il redessine ce que l'on croyait savoir sur ces reptiles marins préhistoriques. Une découverte qui a fait trembler le monde de la paléontologie britannique. 
16/06/2026 futura-terre
Un fossile de « Sword Dragon » vieux de 190 millions d’années réécrit l’histoire des ichtyosaures
Royaume-Uni Jurassique fossile Ichthyosauria évolution squelette
Un ichtyosaure nouvellement identifié de la côte jurassique du Royaume-Uni est en train de réécrire une partie du manuel préhistorique. Surnommé le « dragon-épée du Dorset », ce reptile marin de trois mètres de long a vécu pendant une période d’évolution mal comprise, au cours de laquelle les principaux groupes d’ichtyosaures disparaissaient et de nouveaux apparaissaient. Son squelette magnifiquement préservé – doté d’un museau en forme de lame et d’un éventuel dernier repas – permet de déterminer le moment où cette transition dramatique s’est produite.
24/02/2026 sciencedaily ⚙ Traduction automatique
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