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Photograph of UALVP 24238. Cranial crest and wing phalanges 2-4 of the left wing have been reconstructed with plaster and paint (on the right side of the photograph), as well as metatarsals of the left foot.

Figure 1. Holotype NHMUK PV R 37591 pelvis and vertebrae of P. milnerae gen. et sp. nov. in (a) left lateral view and (b) right lateral view. atr, antitrochanter; bf, brevis fossa; bfr, brevis fossa rim; bs, brevis shelf; dv, dorsal vertebra; iss, ischial shaft; nc, neural canal; no, notch; obf, obturator foramen; poap, postacetabular process; prap, preacetabular process; puf, pubic fenestra; pus, pubic shaft; ras, rib attachment scar; ri, rim; sac, supra-acetabular crest; sv, sacral vertebra.

The holotype (MPSC R 2089) of Aratasaurus museunacionali gen. et sp. nov., photos and drawings from the lateral (A, B) and (C, D) medial views. Abbreviations: cnc, cnemial crest; fe, femur; fosme, fossa medial; lcnrig, lateral cnemial rigde; ti, tibia; tic, tibia crest. Scale bar: 50 mm.

Femur and tibia of Aniksosaurus darwini. Right femur (MDT-PV 1/3) in posterior (A) and medial (B) views. Right tibia (MDT-PV 1/48) in lateral (C) and anterior (D) views. Abbreviations: ast, astragalar depression; at, anterior trochanter; cf, fibular crest ( = crista fibularis); cfl-bf, caudofemoralis longus/brevis fossa; cn, cnemial crest; ec, ectocondylar tuberosity; fc, fibular condyle; fh, femoral head; 4tr, fourth trochanter; gt, greater trochanter; ic, inner condyle; im, inner malleolus; om, outer malleolus; pig, posterior intercondylar groove; tc, tibial condyle; tfc, tibio-fibular crest.

(A) Skeletal reconstruction in left lateral view (missing and damaged portions of the bones in gray). (B) Left ilium in lateral view. (C) Proximal caudal vertebrae in left lateral view with close-up of the infraprezygapophyses. (D) Chevron in cranial view. (E-F) Right scapula in dorsal (E) and lateral (F) views. (G) Last sacral and the two proximalmost caudals in left lateral view. (H) Right pubis in medial view. (I) Right ischium in lateral view. (J) Right femur in distal view. (K) Left metatarsus and distal tarsals in proximal view. (L) Right femur in cranial view. (M-N) Left metatarsus in lateral (M) and dorsal (N) views. Abbreviations: acr, acromion process; ant, antitrochanter; ch, chevron; cv, caudal vertebra(e); diprf, dorsal infraprezygapophyseal fossa; dt, distal tarsal(s); fct, cranial trochanter of femur; fh, femoral head; gl, glenoid fossa; idf, infradiapophyseal fossa; lc, lateral condyle; mc, medial condyle; mep, medial epicondyle; miprf, middle infraprezygapophyseal fossa; mt II, metatarsal II; mt IV, metatarsal IV; mt V, metatarsal V; ns, neural spine; obp, obturator process; pra, preacetabular process; pup, pubic peduncle; sprf, supraprezygapophyseal fossa; sv, sacral vertebra; tfc, tibiofibular crest; tp, transverse process; viprf, ventral infraprezygapophyseal fossa. Scale bars equal 10 cm in (A); 1 cm in (B-N). https://doi.org/10.1371/journal.pone.0210867.g004

Chadititan calvoi nearly complete right femur of MPCN-Pv 1038 in (A) anterior, (B) posterior, (F) proximal, and (G) distal views; distal end of right femur of holotype MPCN-Pv 1034 in (C) anterior, (D) posterior, (E) distal, (H) lateral, and (I) medial views; proximal end of right femur of MPCN-Pv 1035, in (J) proximal, (K), posterior, (L) anterior, and (M) lateral views. Abbreviations: eg, extensor groove; tfc, tibiofibular crest condyle; fg, femoral groove; fh, femoral head; lc, lateral condyle; mc, medial condyle; mt, major trochanter. Scale bar = A-I, 20 cm; J-L, 10 cm.

Fig. 2. Right tibia of Quilmesaurus curriei compared to other tibiae of Abelisauridae. A-B: Quilmesaurus curriei in lateral and anterior view; C: Aucasaurus garridoi, D: Genusaurus sisterornis, E and H: Ekrixinatosaurus novasi, F: Indosuchus raptorius (Courtesy of Dr. F. E. Novas), G: Rajasaurus narmadensis; I: Pycnonemosaurus nevesi Abbreviations: faap: facet for the ascending process of the astragalus; cc: cnemial crest; lm: lateral malleolus; lma: lateral margin of the malleolus; lpcc: lateral process of the cnemial crest; mm: medial malleolus; mma: medial margin of the malleolus; vmcc: ventral margin of the cnemial crest; vpcc: ventral process of the cnemial crest.

Fig. 2. Right tibia of Quilmesaurus curriei compared to other tibiae of Abelisauridae. A-B: Quilmesaurus curriei in lateral and anterior view; C: Aucasaurus garridoi, D: Genusaurus sisterornis, E and H: Ekrixinatosaurus novasi, F: Indosuchus raptorius (Courtesy of Dr. F. E. Novas), G: Rajasaurus narmadensis; I: Pycnonemosaurus nevesi Abbreviations: faap: facet for the ascending process of the astragalus; cc: cnemial crest; lm: lateral malleolus; lma: lateral margin of the malleolus; lpcc: lateral process of the cnemial crest; mm: medial malleolus; mma: medial margin of the malleolus; vmcc: ventral margin of the cnemial crest; vpcc: ventral process of the cnemial crest.

Lusotitan atalaiensis. Photographs of right humerus (proximal half) in (A) anterior (slightly oblique as a result of mounted position), (B) medial, (C) proximal, (D) lateral, and (E) posterior views. Abbreviations: dtp, deltopectoral crest; hh, humeral head. Scale bar = 200 mm.

Eopteranodon lii holotype (BPV 078). (A) Counterpart; (B) main part. (C and D) Respective schematic drawings. Abbreviations: cv, cervical vertebra; co, coracoid; d1–d4, digits 1–4; fe, femur; fi, fibula; h, humerus; j, jugal; mand, mandible; mc, metacarpal; pmc, premaxillary crest; pe, pelvis; ph, phalanx; ti, tibia; ul, ulna; rad, radius. Scale bars: C, 50 mm; D, 10 mm.

Mimodactylus libanensis gen. et sp. nov. (a) Photo and drawing of the complete specimen. (b) Close up of scapula and coracoid. (c) Detail of the wrist, showing the relation of the pteroid and the carpus. (d) Detail of the humerus. Scale-bars, a: 50 mm; b-d: 10 mm. Abbreviations. car: carpus; cdv: caudal vertebrae; cor: coracoid; cra: cranium; cs: cristospine; cv: cervical vertebrae; d: dentary; dcar: distal carpals; dpc: deltopectoral crest; dri: dorsal ribs; dv: dorsal vertebrae; fe: femur; hu: humerus; man: mandible; mcI-III: first to third metacarpals; mcIV: wing metacarpal; mtar: metatarsals; pcar: proximal carpals; ph1d4: first wing phalanx; ph2d4: second wing phalanx; ph3d4: third wing phalanx; ph4d4: fourth wing phalanx; ppu: prepubis; ptd: pteroid; ra: radius; sca: scapula; sri: sacral ribs; stp: sternal plate; sym: mandibular symphysis; tar: tarsus; te: teeth; ti: tibia; ul: ulna; the abbreviations ‘l’ and ‘r’ represents respectively left and right.

Mimodactylus libanensis gen. et sp. nov. (a) Photo and drawing of the complete specimen. (b) Close up of scapula and coracoid. (c) Detail of the wrist, showing the relation of the pteroid and the carpus. (d) Detail of the humerus. Scale-bars, a: 50 mm; b-d: 10 mm. Abbreviations. car: carpus; cdv: caudal vertebrae; cor: coracoid; cra: cranium; cs: cristospine; cv: cervical vertebrae; d: dentary; dcar: distal carpals; dpc: deltopectoral crest; dri: dorsal ribs; dv: dorsal vertebrae; fe: femur; hu: humerus; man: mandible; mcI-III: first to third metacarpals; mcIV: wing metacarpal; mtar: metatarsals; pcar: proximal carpals; ph1d4: first wing phalanx; ph2d4: second wing phalanx; ph3d4: third wing phalanx; ph4d4: fourth wing phalanx; ppu: prepubis; ptd: pteroid; ra: radius; sca: scapula; sri: sacral ribs; stp: sternal plate; sym: mandibular symphysis; tar: tarsus; te: teeth; ti: tibia; ul: ulna; the abbreviations ‘l’ and ‘r’ represents respectively left and right.

Cimoliopterus cuvieri. Holotype NHMUK PV 39409 (Cenomanian / Turonian, Chalk Formation), anterior part of the rostrum A right lateral view B respective line drawing C ventral view D respective line drawing. Abbreviations: m – maxillae, pm – premaxillae, pmcr – premaxillary crest, prid – palatal ridge. Arrows and numbers indicate alveoli or teeth and their respective position. Scale bar = 10 mm. Photos courtesy of The Natural History Museum.

T. regalis FSAC-OB 1, holotype left humerus. In (A), ventral view, (B) dorsal view, (C) anterior view, (D) distal view, and (E) proximal view. Abbreviations: dpc, deltopectoral crest; ect, ectepicondyle; ent, entepicondyle; lc, lateral condyle; mc, medial condyle; pf, pneumatic fossa/foramen; scpr, supracondylar process; uc, ulnar crest; ut, ulnar tubercle.

Lonchodraco giganteus comb. n. Lectotype NHMUK PV 39412 (Cenomanian / Turonian, Chalk Formation). A–F articulated anterior parts of the rostrum and mandible A right lateral view B respective line drawing C left lateral view D respective line drawing E anterior view F respective line drawing G associated scapulocoracoid in posterior view. Abbreviations: ch – choanae, cor – coracoid, d – dentary, dcr – dentary crest, m – maxillae, pl – palatine, pm – premaxillae, pmcr – premaxillaery crest, prid – palatal ridge, sca – scapula. Arrows indicate alveoli or teeth. Scale bar = 10 mm. Photos courtesy of The Natural History Museum.

Plesiolophus is a lambeosaurine hadrosaur known from the same locality, but an earlier time period than Parasaurolophus. Though it is nearly impossible to know conclusively, Plesiolophus cannot be ruled out as a direct ancestor. Like Parasaurolophus, Plesiolophus would have had a distinct crest protruding backward from its skull.