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Representative taxa from the Santonian Iharkút fauna from the Csehbánya Formation, Bakony Mountains, western Hungary. A Pannoniasaurus inexpectatus (Squamata, Mosasauroidea), dorsal vertebra (MTM uncatalogued) in dorsal view (photo by Réka Kalmár) B Foxemys trabanti (Pleurodira, Bothremydidae), skull (MTM V 2010.215.1.) in dorsal view (photo by Márton Rabi). C Bicuspidon aff. hatzegiensis (Squamata, Borioteiioidea), left dentary (MTM 2006.112.1.) in medial view (photo by László Makádi) D Basal tetanuran (Theropoda, Tetanurae), tooth (MTM V.01.54) in ?lingual view E Indeterminate abelisaurid (Theropoda, Abelisauridae), pedal ungual phalanx (MTM V 2008.43.1.) in lateral view F Pneumatoraptor fodori (Theropoda, Paraves), left scapulocoracoid (holotype, MTM V 2008.38.1.) in lateral view G Mochlodon vorosi (Ornithopoda, Rhabdodontidae), left dentary (holotype, MTM V 2010.105.1) in lateral view H Bakonydraco galaczi (Pterosauria, Azhdarchidae), mandible (holotype, MTM 2007.110.1) in dorsal view I Iharkutosuchus makadii (Eusuchia, Hylaeochampsidae), skull (holotype, MTM 2006.52.1) in dorsal view J Hungarosaurus tormai (Ankylosauria, Nodosauridae), right dentary (MTM 2007.25.2) in lateral view K Bauxitornis mindszentyae (Aves, Enantiornithes), left tarsometatarsus (holotype, MTM V 2009.38.1) in anterior view L Ajkaceratops kozmai (Ceratopsia), fused rostral and premaxillae (holotype, MTM V 2009.192.1) in lateral view. Scale bars: 2 cm in A, V, G, H, I, J; 1 cm in D, E, F, K, L; 1 mm in C.

Representative taxa from the Santonian Iharkút fauna from the Csehbánya Formation, Bakony Mountains, western Hungary. A Pannoniasaurus inexpectatus (Squamata, Mosasauroidea), dorsal vertebra (MTM uncatalogued) in dorsal view (photo by Réka Kalmár) B Foxemys trabanti (Pleurodira, Bothremydidae), skull (MTM V 2010.215.1.) in dorsal view (photo by Márton Rabi). C Bicuspidon aff. hatzegiensis (Squamata, Borioteiioidea), left dentary (MTM 2006.112.1.) in medial view (photo by László Makádi) D Basal tetanuran (Theropoda, Tetanurae), tooth (MTM V.01.54) in ?lingual view E Indeterminate abelisaurid (Theropoda, Abelisauridae), pedal ungual phalanx (MTM V 2008.43.1.) in lateral view F Pneumatoraptor fodori (Theropoda, Paraves), left scapulocoracoid (holotype, MTM V 2008.38.1.) in lateral view G Mochlodon vorosi (Ornithopoda, Rhabdodontidae), left dentary (holotype, MTM V 2010.105.1) in lateral view H Bakonydraco galaczi (Pterosauria, Azhdarchidae), mandible (holotype, MTM 2007.110.1) in dorsal view I Iharkutosuchus makadii (Eusuchia, Hylaeochampsidae), skull (holotype, MTM 2006.52.1) in dorsal view J Hungarosaurus tormai (Ankylosauria, Nodosauridae), right dentary (MTM 2007.25.2) in lateral view K Bauxitornis mindszentyae (Aves, Enantiornithes), left tarsometatarsus (holotype, MTM V 2009.38.1) in anterior view L Ajkaceratops kozmai (Ceratopsia), fused rostral and premaxillae (holotype, MTM V 2009.192.1) in lateral view. Scale bars: 2 cm in A, V, G, H, I, J; 1 cm in D, E, F, K, L; 1 mm in C.

tooth vertebra Hungary Santonian +14
Representative taxa from the latest Campanian–Maastrichtian faunas from Transylvania, western Romania. A–B Nidophis insularis (Serpentes, Madtsoiidae), articulated vertebrae (LPB (FGGUB) v.547/2) in left lateral (A) and dorsal (B) views (Densuş-Ciula Formation, Tuştea, Haţeg Basin; photo by Ştefan Vasile) C Allodaposuchus precedens (Eusuchia, ?Hylaeochampsidae), skull (PSMUBB V 438) in dorsal view (Sebeş = Şard Formation, Oarda de Jos, southwestern Transylvanian Basin; photo by Vlad Codrea/Massimo Delfino) D Theriosuchus sympiestodon (Mesoeucrocodylia, Atoposauridae), right maxilla (MCDRD 793) in lateral view (Sînpetru Formation, Sînpetru, Haţeg Basin) E–F Indeterminate titanosaur (?Magyarosaurus dacus) (Sauropoda, Titanosauria), isolated osteoderm (LPB (FGGUB) R.1410) in dorsal (E) and lateral (F) views (Sînpetru Formation, Sînpetru, Haţeg Basin) G Indeterminate ornithuran bird (Aves, Ornithurae), incomplete left tibiotarsus (LPB (FGGUB) R.1902) in anterior view (Densuş-Ciula Formation, Vălioara, Haţeg Basin) H Balaur bondoc (Theropoda, Dromaeosauridae), articulated left distal hindlimb (EME PV.313) in lateral view (Sebeş = Şard Formation, Sebeş-Glod, southewestern Transylvanian Basin; photo by Mick Ellison) I Zalmoxes robustus (Ornithopoda, Rhabdodontidae), right dentary (NHMUK R.3407) in medial view (Sînpetru Formation, Sînpetru, Haţeg Basin) J Telmatosaurus transsylvanicus (Hadrosauria), right maxilla (MFGI unnumbered) in lateral view (Sînpetru Formation, Sînpetru, Haţeg Basin) K Indeterminate nodosaurid – Struthiosaurus transylvanicus or new taxon – (Ankylosauria, Nodosauridae), isolated tooth (LPB (FGGUB) R.2182) in medial view (Sînpetru Formation, Sînpetru, Haţeg Basin) L Barbatodon transylvanicus (Multituberculata, Kogaionidae), right maxilla (LPB (FGGUB) M.1635) in medial view (Sînpetru Formation, Pui, Haţeg Basin). Scale bars equal 1 mm in A, B; 5 mm in K; 1 cm in G, L; 2 cm in D; and 5 cm in C, E, F, H, I, J.

Representative taxa from the latest Campanian–Maastrichtian faunas from Transylvania, western Romania. A–B Nidophis insularis (Serpentes, Madtsoiidae), articulated vertebrae (LPB (FGGUB) v.547/2) in left lateral (A) and dorsal (B) views (Densuş-Ciula Formation, Tuştea, Haţeg Basin; photo by Ştefan Vasile) C Allodaposuchus precedens (Eusuchia, ?Hylaeochampsidae), skull (PSMUBB V 438) in dorsal view (Sebeş = Şard Formation, Oarda de Jos, southwestern Transylvanian Basin; photo by Vlad Codrea/Massimo Delfino) D Theriosuchus sympiestodon (Mesoeucrocodylia, Atoposauridae), right maxilla (MCDRD 793) in lateral view (Sînpetru Formation, Sînpetru, Haţeg Basin) E–F Indeterminate titanosaur (?Magyarosaurus dacus) (Sauropoda, Titanosauria), isolated osteoderm (LPB (FGGUB) R.1410) in dorsal (E) and lateral (F) views (Sînpetru Formation, Sînpetru, Haţeg Basin) G Indeterminate ornithuran bird (Aves, Ornithurae), incomplete left tibiotarsus (LPB (FGGUB) R.1902) in anterior view (Densuş-Ciula Formation, Vălioara, Haţeg Basin) H Balaur bondoc (Theropoda, Dromaeosauridae), articulated left distal hindlimb (EME PV.313) in lateral view (Sebeş = Şard Formation, Sebeş-Glod, southewestern Transylvanian Basin; photo by Mick Ellison) I Zalmoxes robustus (Ornithopoda, Rhabdodontidae), right dentary (NHMUK R.3407) in medial view (Sînpetru Formation, Sînpetru, Haţeg Basin) J Telmatosaurus transsylvanicus (Hadrosauria), right maxilla (MFGI unnumbered) in lateral view (Sînpetru Formation, Sînpetru, Haţeg Basin) K Indeterminate nodosaurid – Struthiosaurus transylvanicus or new taxon – (Ankylosauria, Nodosauridae), isolated tooth (LPB (FGGUB) R.2182) in medial view (Sînpetru Formation, Sînpetru, Haţeg Basin) L Barbatodon transylvanicus (Multituberculata, Kogaionidae), right maxilla (LPB (FGGUB) M.1635) in medial view (Sînpetru Formation, Pui, Haţeg Basin). Scale bars equal 1 mm in A, B; 5 mm in K; 1 cm in G, L; 2 cm in D; and 5 cm in C, E, F, H, I, J.

Romania Densuş-Ciula Sebeş Sînpetru +9
Rates of skeletal character evolution in the skull and postcranial skeleton of hadrosauroids. Cladograms illustrate the results from branch likelihood tests for two morphological partitions: skull (cranium and mandible) (A) and postcranial skeleton (B). In both cladograms, results from the branch likelihood tests are summarized on a strict consensus tree derived from four separately analyzed MPTs, each with 100 dating replicates (a total of 400 Hedman-dated phylogenies). Pie charts on branches illustrate the proportion of dating replicates that showed significantly high rates (red), slow rates (blue), or nonsignificant average rates (white). No pie charts are plotted on branches that showed nonsignificant rates in 100% of dating replicates. Branches that showed high rates (red) in more than 50% of dating replicates are doubled in length. See the Supplementary Material for Hedman-based results plotted separately for each MPT (Supplementary Fig. S2) and for results using the MBL dating method (Supplementary Fig. S3). Silhouettes were created by Scott Hartman and were downloaded from http://phylopic.org (Creative Commons license CC BY 3.0).

Rates of skeletal character evolution in the skull and postcranial skeleton of hadrosauroids. Cladograms illustrate the results from branch likelihood tests for two morphological partitions: skull (cranium and mandible) (A) and postcranial skeleton (B). In both cladograms, results from the branch likelihood tests are summarized on a strict consensus tree derived from four separately analyzed MPTs, each with 100 dating replicates (a total of 400 Hedman-dated phylogenies). Pie charts on branches illustrate the proportion of dating replicates that showed significantly high rates (red), slow rates (blue), or nonsignificant average rates (white). No pie charts are plotted on branches that showed nonsignificant rates in 100% of dating replicates. Branches that showed high rates (red) in more than 50% of dating replicates are doubled in length. See the Supplementary Material for Hedman-based results plotted separately for each MPT (Supplementary Fig. S2) and for results using the MBL dating method (Supplementary Fig. S3). Silhouettes were created by Scott Hartman and were downloaded from http://phylopic.org (Creative Commons license CC BY 3.0).

Protohadros dating evolution skeleton +1
Figure 6. Hadrosaurs of North America, exemplied by the selection of skulls of valid nominal genera. The grey shadows on the skulls represent the reconstructed parts. Highlighted names indicate the unique genera for each region

Figure 6. Hadrosaurs of North America, exemplied by the selection of skulls of valid nominal genera. The grey shadows on the skulls represent the reconstructed parts. Highlighted names indicate the unique genera for each region

Hadrosauria Protohadros skull
Brachylophosaurus canadensis skull (original). From the Oldman Formation, Milk River, Alberta. On display at the Royal Tyrrell Museum, Alberta, Canada.

Brachylophosaurus canadensis skull (original). From the Oldman Formation, Milk River, Alberta. On display at the Royal Tyrrell Museum, Alberta, Canada.

museum Canada Milk River Oldman +3
Holotype of Khunnuchelys lophorhothon sp. nov. from Baybishe, northeastern Aral Sea region, Kazakhstan, Bostobe Formation, Santonian–early Campanian, Late Cretaceous, ZIN PH 5/55, a partial skull; in dorsal (A), ventral (B), lateral (C), and anterior (D) views. Photographs (A1–D1), explanatory drawings (A2–D2).
Formations Bostobe

Holotype of Khunnuchelys lophorhothon sp. nov. from Baybishe, northeastern Aral Sea region, Kazakhstan, Bostobe Formation, Santonian–early Campanian, Late Cretaceous, ZIN PH 5/55, a partial skull; in dorsal (A), ventral (B), lateral (C), and anterior (D) views. Photographs (A1–D1), explanatory drawings (A2–D2).

drawing Kazakhstan Bostobe Campanian +8
Restoration of the spinosaurid dinosaur Siamosaurus in the Sao Khua Formation palaeoenvironment, with Sunosuchus in the middle left and a herd of Phuwiangosaurus in the background.
References:
Siamosaurus based on tooth specimens [1] and the neural spine of a possibly referable skeleton[2], with other missing elements filled in with relatives (Suchomimus[3], Baryonyx[4], IchthyovenatorFile:Ichthyovenator_laosensis_skeletal_reconstruction_by_PaleoGeek.png).
Phuwiangosaurus based on skeletal by Suteethorn et al. (2009)[5] and missing elements of skull of EuhelopusFile:Euhelopus.png.

Sunosuchus based on Suteethorn and Ingavat (1983)[6] and missing elements based on Goniopholis[7].

Restoration of the spinosaurid dinosaur Siamosaurus in the Sao Khua Formation palaeoenvironment, with Sunosuchus in the middle left and a herd of Phuwiangosaurus in the background. References: Siamosaurus based on tooth specimens [1] and the neural spine of a possibly referable skeleton[2], with other missing elements filled in with relatives (Suchomimus[3], Baryonyx[4], IchthyovenatorFile:Ichthyovenator_laosensis_skeletal_reconstruction_by_PaleoGeek.png). Phuwiangosaurus based on skeletal by Suteethorn et al. (2009)[5] and missing elements of skull of EuhelopusFile:Euhelopus.png. Sunosuchus based on Suteethorn and Ingavat (1983)[6] and missing elements based on Goniopholis[7].

tooth drawing Sao Khua specimen +11
Juvenile Gorgosaurus TMP 2009.12.14 preserving stomach contents. Photographs of specimen in (A) right lateral view and (B) left anterolateral view. (C) Interpretive illustration of specimen in right lateral view. Skeleton consists of a nearly complete skull, the left side of the body and limbs, and a nearly complete pelvis. Red rectangle delineates location of stomach contents. (D) Histological photomicrograph of tibia showing the presence of five lines of arrested growths and two annuli (marked by asterisks), indicating that the individual was between 5 and 7 years old. Scale bars, 50 cm (A) to (C) and 1 mm (D).

Juvenile Gorgosaurus TMP 2009.12.14 preserving stomach contents. Photographs of specimen in (A) right lateral view and (B) left anterolateral view. (C) Interpretive illustration of specimen in right lateral view. Skeleton consists of a nearly complete skull, the left side of the body and limbs, and a nearly complete pelvis. Red rectangle delineates location of stomach contents. (D) Histological photomicrograph of tibia showing the presence of five lines of arrested growths and two annuli (marked by asterisks), indicating that the individual was between 5 and 7 years old. Scale bars, 50 cm (A) to (C) and 1 mm (D).

limb pelvis scale growth +6
Coelophysidae skull restoration

Coelophysidae skull restoration

Coelophysidae Syntarsus skull
Anhanguera blittersdorfi skull

Anhanguera blittersdorfi skull

Anhanguera skull
Skull of Platecarpus coryphaeus

Skull of Platecarpus coryphaeus

Platecarpus skull
Complete specimen, excellent mineralization. Presented with a base (exceptional, very good condition)).	
Skull length 18,9 inch ( 28,3 inch with vertebras)

Complete specimen, excellent mineralization. Presented with a base (exceptional, very good condition)). Skull length 18,9 inch ( 28,3 inch with vertebras)

vertebra specimen Platecarpus skull
Crâne de mosasaure d'espèce Platecarpus tympaniticus (squamates, mosasaures).
Provenance : Smoky Hill Chalk, Kansas (aux Etats-Unis).
Date : Crétacé supérieur, période du Campanien, 88 millions d'années avant notre ère.
Collections du Muséum national d'histoire naturelle de Paris (France).
Exposé à l'occasion de l'exposition "Un T-Rex à Paris" au Muséum national d'histoire naturelle de Paris (France) du 6 juin au 2 septembre 2018.

Légende du fossile dans cette exposition : "Les mosasaures ne sont pas des dinosaures mais des reptiles marins, très répandus à la fin du Crétacé. Certains genres comme Globidens et Halisaurus sont connus aussi bien aux Etats-Unis qu'au Maroc. Ces animaux étaient d'excellents nageurs, capables de traverser l'Atlantique."

Crâne de mosasaure d'espèce Platecarpus tympaniticus (squamates, mosasaures). Provenance : Smoky Hill Chalk, Kansas (aux Etats-Unis). Date : Crétacé supérieur, période du Campanien, 88 millions d'années avant notre ère. Collections du Muséum national d'histoire naturelle de Paris (France). Exposé à l'occasion de l'exposition "Un T-Rex à Paris" au Muséum national d'histoire naturelle de Paris (France) du 6 juin au 2 septembre 2018. Légende du fossile dans cette exposition : "Les mosasaures ne sont pas des dinosaures mais des reptiles marins, très répandus à la fin du Crétacé. Certains genres comme Globidens et Halisaurus sont connus aussi bien aux Etats-Unis qu'au Maroc. Ces animaux étaient d'excellents nageurs, capables de traverser l'Atlantique."

France Morocco Campanian Cretaceous +8
Drawing of the skull of MCZ 4374, the holotype of Macrosaurus proriger (Tylosaurus proriger) from Cope (1870)

Drawing of the skull of MCZ 4374, the holotype of Macrosaurus proriger (Tylosaurus proriger) from Cope (1870)

drawing holotype Drinker Hainosaurus +3
a Skeletal reconstruction of A. greppini. Elements preserved in the material and therefore providing information for the skeletal reconstruction are marked in blue. Because much information is missing from the incomplete skeletal material, the dorsal vertebrae, the proportions and morphology of the cervical vertebrae and the skull were modified from Camarasaurus. b Scaled silhouette drawings of Cetiosauriscus stewarti (in black) and A. greppini (in grey) demonstrating the significant size difference between the two taxa. Scale bar is 1 m

a Skeletal reconstruction of A. greppini. Elements preserved in the material and therefore providing information for the skeletal reconstruction are marked in blue. Because much information is missing from the incomplete skeletal material, the dorsal vertebrae, the proportions and morphology of the cervical vertebrae and the skull were modified from Camarasaurus. b Scaled silhouette drawings of Cetiosauriscus stewarti (in black) and A. greppini (in grey) demonstrating the significant size difference between the two taxa. Scale bar is 1 m

scale drawing Amanzia Camarasaurus +2
Reconstructed skull of the turiasaurian Mierasaurus, based on the holotype UMNH.VP.26004.

Reconstructed skull of the turiasaurian Mierasaurus, based on the holotype UMNH.VP.26004.

holotype Mierasaurus Turiasauria skull
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News

A 250-million-year-old fossil reveals the origins of mammal hearing
bone jaw fossil mammals skull
Sensitive hearing may have evolved in mammal ancestors far earlier than scientists once believed. By modeling how sound moved through the skull of Thrinaxodon, a 250-million-year-old mammal predecessor, researchers found it likely used an early eardrum to hear airborne sounds. This challenges the long-held idea that these animals mainly “listened” through their jaws or bones. The results reveal that a key feature of modern mammal hearing was already taking shape deep in prehistory.
20/01/2026 sciencedaily
Anurognathus: Beast of the Week
Anurognathus: Beast of the Week
Germany Jurassic Late Jurassic Anurognathus Pterosauria skull
This week we will be checking out a unique little pterosaur, Anurognathus ammoni!  Anurognathus lived in what is now Germany during the late Jurassic period, about 150 million years ago.  It was tiny, sporting a 14 inch (35.5 cm) wingspan, and would have likely eaten insects.  It's genus name translates to "Frog Jaw" since its skull was similar looking to a frog's, being extremely blunt with a wide mouth.  Watercolor reconstruction of Anurognathus ammoni by Christopher DiPiazza.Anurognathus' sku
14/12/2025 prehistoricbeastoftheweek
242-million-year-old mini predator changes lizard evolution
limb tooth predator fossil evolution new species skull
A tiny 242-million-year-old fossil from Devon is shaking up scientists’ assumptions about the earliest members of the lizard lineage. Instead of the expected skull hinges and palate teeth typical of modern lizards and snakes, this ancient creature shows a surprising mix of primitive and unusual traits—along with strikingly large, blade-like teeth. High-resolution synchrotron scans revealed details invisible to the naked eye, helping researchers name the new species Agriodontosaurus helsbypetrae
30/11/2025 sciencedaily
Humans evolved faster than any other ape
growth evolution skull
UCL scientists found that human skulls evolved much faster than those of other apes, reflecting the powerful forces driving our brain growth and facial flattening. By comparing 3D models of ape skulls, they showed that humans changed about twice as much as expected. The findings suggest that both cognitive and social factors, not just intelligence, influenced our evolutionary path.
29/10/2025 sciencedaily-human-evo
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