Campanien

Elasmosaurus platyurus, plesiosaurian from Late Cretaceous (Campanian) of North America

Elasmosaurus platyurus, plesiosaurian from Late Cretaceous (Campanian) of North America

Latoplatecarpus willistoni (TMP 1984.162.0001), Royal Tyrrell Museum, Drumheller, Alberta, 2025-07-13; cast, Campanian, from the Pierre Shale

Aralazhdarcho bostobensis, ZIN PH 57/43, a proximal fragment of a left humerus in proximal (a), ventral (b), anterior (c), dorsal (d), and posterior (e) views. This specimen is from the Shakh Shakh II locality in the northeasten Aral Sea region of Kazakhstan; Bostobe Formation, Upper Cretaceous (Santonian – lower Campanian). Abbreviations: h, humeral head; pf, pneumatic foramen; uc, ulnar crest. Scale bar is 10 mm.

Plate 53. Borogovia gracilicrus gen. et sp. n. 1. Fragmentary right pes, in: a dorsal, b ventral, views; ZPAL MgD-11174, holotype, X 0.5. 2. Phalanx 11-1 of the left pes, medial view; same specimen, X 1. 3. Ungual of left second pedal digit, ventral view; same specimen, X 1. 4. Phalanx 111-3 of the left pes, lateral view; same specimen, X 1. 5. Phalanges IV-1 to IV-4 of the left pes, lateral view; same specimen, X 1. 6. Distal portion of right metatarsal 111, posterior view; same specimen, X 1. 7. Distal portion of left metatarsal IV, medial view; same specimen, X 1. 8. Distal portion of left metatarsal 11, lateral view; same specimen, X 1. 9. Right tibiotarsus, a distal portion and b proximal portion with fragment of the fibula attached, posterior views, c distal portion, anterior view, d distal portion, lateral view; same specimen, X 0.5. Nemegt Formation, ?Late Campanian or ?Early Maastrichtian, Ultan Ula IV, Nemegt Basin, Gobi Desert, Mongolia

Skull of troodontid theropod Xixiasaurus henanensis gen. et sp. nov. (HGM 41HIII−0201; holotype), lower–middle Majiacun Formation (Upper Cretaceous: Coniacian–Campanian), Henan Province, China; in dorsal (A); lateral (B), and ventral (C) views. Note that this version of the image does not include the interpretative lines of the version in the paper.

Reconstruction of the skull of Spinops sternbergorum gen. et sp. nov. from the Campanian of Dinosaur Provincial Park, southern Alberta, in right lateral view. Preserved elements are stippled; missing portions are dotted and modeled after Centrosaurus apertus.

Photograph of the holotype skull of Titanoceratops ouranos, from the late Campanian aged Kirtland/Fruitland beds of New Mexico, USA. Photograph by Nick Longrich

Representative taxa from the early Campanian Muthmannsdorf fauna from the Grünbach Formation, eastern Austria. A Doratodon carcharidens (Mesoeucrocodylia) mandible (PIUW 2349/57) in dorsal view (photo by Márton Rabi) B Indeterminate azhdarchid (Pterosauria, Azhdarchidae), left humerus (PIUW 2349/102) in anterior view C ‘Megalosaurus pannoniensis’ basal tetanuran (Theropoda, Tetanurae), tooth (PIUW uncatalogued) in lateral view D Mochlodon suessi (Ornithopoda, Rhabdodontidae), right dentary (holotype, PIUW 2349/2) in medial view. Scale bars equal 2 cm in A, B and D and 1 cm in C.

Representative taxa from the latest Campanian–Maastrichtian faunas from Transylvania, western Romania. A–B Nidophis insularis (Serpentes, Madtsoiidae), articulated vertebrae (LPB (FGGUB) v.547/2) in left lateral (A) and dorsal (B) views (Densuş-Ciula Formation, Tuştea, Haţeg Basin; photo by Ştefan Vasile) C Allodaposuchus precedens (Eusuchia, ?Hylaeochampsidae), skull (PSMUBB V 438) in dorsal view (Sebeş = Şard Formation, Oarda de Jos, southwestern Transylvanian Basin; photo by Vlad Codrea/Massimo Delfino) D Theriosuchus sympiestodon (Mesoeucrocodylia, Atoposauridae), right maxilla (MCDRD 793) in lateral view (Sînpetru Formation, Sînpetru, Haţeg Basin) E–F Indeterminate titanosaur (?Magyarosaurus dacus) (Sauropoda, Titanosauria), isolated osteoderm (LPB (FGGUB) R.1410) in dorsal (E) and lateral (F) views (Sînpetru Formation, Sînpetru, Haţeg Basin) G Indeterminate ornithuran bird (Aves, Ornithurae), incomplete left tibiotarsus (LPB (FGGUB) R.1902) in anterior view (Densuş-Ciula Formation, Vălioara, Haţeg Basin) H Balaur bondoc (Theropoda, Dromaeosauridae), articulated left distal hindlimb (EME PV.313) in lateral view (Sebeş = Şard Formation, Sebeş-Glod, southewestern Transylvanian Basin; photo by Mick Ellison) I Zalmoxes robustus (Ornithopoda, Rhabdodontidae), right dentary (NHMUK R.3407) in medial view (Sînpetru Formation, Sînpetru, Haţeg Basin) J Telmatosaurus transsylvanicus (Hadrosauria), right maxilla (MFGI unnumbered) in lateral view (Sînpetru Formation, Sînpetru, Haţeg Basin) K Indeterminate nodosaurid – Struthiosaurus transylvanicus or new taxon – (Ankylosauria, Nodosauridae), isolated tooth (LPB (FGGUB) R.2182) in medial view (Sînpetru Formation, Sînpetru, Haţeg Basin) L Barbatodon transylvanicus (Multituberculata, Kogaionidae), right maxilla (LPB (FGGUB) M.1635) in medial view (Sînpetru Formation, Pui, Haţeg Basin). Scale bars equal 1 mm in A, B; 5 mm in K; 1 cm in G, L; 2 cm in D; and 5 cm in C, E, F, H, I, J.

Campanian fauna from the Oldman Formation: Wendiceratops, Corythosaurus, Scolosaurus, Chasmosaurus and "Troodon"

Brachylophosaurus canadensis (TMP 2005.000.0029), Royal Tyrrell Museum, Drumheller, Alberta, 2025-07-13; cast, Campanian, from the Oldman Formation

African locality: 5, Majunga, Maevarano Formation (Campanian): Rahonavis ostromi.

Locality map: Grand Staircase-Escalante National Monument, southern Utah. Map showing the Nipple Butte area (indicated by yellow star) of Grand Staircase-Escalante National Monument (GSENM). GSENM is bounded by the red rectangle and silhouetted in dark gray on the inset of Utah and surrounding states (modified from [1]). The original map has been modified to show the Nipple Butte area instead of the Machairoceratops fossil locality as in the original source. New location based on map in A New Macrovertebrate Assemblage from the Late Cretaceous (Campanian) of Southern Utah, page 601.

Kaiparowits Formation (Campanian; Upper Cretaceous) at The Blues, near Powell Point (peak to the left) composed of the Claron Formation.

Map of Grand Staircase-Escalante National Monument (GSENM), southern Utah (A), and generalized stratigraphic section of the upper Campanian Kaiprowits Formation (B). The approximate stratigraphic position of Akainacephalus johnsoni is located near the base of the middle unit within the Kaiparowits Formation. The map highlights the GSENM boundary (dashed line), showing the geological distribution and outcrops of the Cretaceous Kaiparowits, Wahweap, and Straight Cliffs formations. The red star indicates the Horse Mountain area, from which Akainacephalus johnsoni was recorded. Map and stratigraphic column modified from Roberts (2005). Radioisotopic dates used from Roberts, Deino & Chan (2005) and Roberts et al. (2013), respectively.