Dinosauria

Map of the localities in the Bauru Basin where the sauropod dinosaurs were collected.

(A) A phylogenetic principal-component analysis (PCA) represents the projection of the Dinosauria supertree (STAR Methods) into a PCA of climatic variables. PC1 axis shows strong positive correlation with maximum temperature ([temp max), low positive correlation with precipitation seasonality ( precip season), strong negative correlation with minimum temperature (Ytemp min), and strong negative correlation with minimum precipitation (Yprecip min). PC2 axis shows strong positive correlation with minimum temperature ([temp min) and negative correlation with precipitation seasonality (Yprecip season). Shadows around points highlight the relative density in the principal compo- nent space of non-dinosaurian Dinosauromorpha (gray), Ornithischia (blue), Sauropodomorpha (green), and Theropoda (red). (B) Lower left plot shows 95% confidence interval convex hulls for each dinosauromorph subclade. Blue thermometer (top left corner) symbolizes the direction of the vector in the PC space region for cold temper- atures; yellow thermometer (top right corner) indicates the direction of the vector in PC space for warm tem- peratures; brown shrub (top right corner) depicts dry conditions, while the same with a gray, rainy cloud (mid, lower side of the graph) illustrates seasonal conditions. Silhouettes represent the following taxa (clockwise from the higher left corner): Minmi, Edmontosaurus, Pachyrhinosaurus, Tyrannosaurus, Asilisaurus, Graci- liceratops, Harpymimus, Altirhinus, Gobititan, Suz- housaurus, Marasuchus, Pampadromaeus, Herrer- asaurus, Vulcanodon, Diplodocus, Giraffatitan, Coelophysis, Dromomeron, Gondwanatitan, Tapuiasaurus, Anchisaurus, Siamotyrannus, Diodorus, Suchomimus, Phuwiangosaurus, Ouranosaurus, Irritator, Tangvayosaurus, Nanshiungosaurus, Aeolosaurus, Rebbachisaurus, Chuxiongosaurus, Tethyshadros, Koreanosaurus. Genyodectes, Mapusaurus, Vegavis, Goyocephale, and Rhoetosaurus.

Fossil samples – e.g. ‘primitive’ bony fish (1, 2), a skull of a temnospondyl ‘amphibian’ (probably a metoposauroid) in dorsal view (3), a skull of an archosaur of the crocodile lineage (probably a phytosaur) in palatal view (4), holotype of the “gliding reptile” Icarosaurus siefkeri [1] (5) and Atreipus-Grallator-type dinosaur tracks (bottom right) – from the Newark Supergroup, i.e. a series of mainly Late Triassic to Early Jurassic sedimentary rocks of eastern North America ↑ Edwin H. Colbert: A gliding reptile from the Triassic of New Jersey. American Museum Novitates, 2230. American Museum of Natural History, New York 1966, digitallibrary.amnh.org, cf. fig. 3 therein.

Diuqin is an unenlagiine dinosaur from the Late Cretaceous of what is now Argentina. Unenlagiines, known exclusively from South America, are usually classified as dromaeosaurs though this is sometimes debated. Like dromaeosaurs, they were covered in feathers, carnivorous, and had the large sickle-like claw on the inner toe of each foot. Unique to unenlagiines is their elongated snout, suggesting a piscivorous diet. Diuqin was a medium-sized unenlagiine, at about 4 m in length.

Figure 1. Evolution of macroecological traits in Dinosauria. Large scale event in dinosaur evolution (a); the origin of dinosaurs (star), hyperthermals (volcano), the earliest fossil Avialae (bird), the earliest fossil angiosperm (flower), the Cretaceous/Palaeogene mass extinction (asteroid). Phylogeny of dinosaurs (b) redrawn from Sereno and adapted to the current consensus and upon which an ancestral state reconstruction of temperature niche (mean annual temperature) after Chiarenza et al. is plotted; Mesozoic palaeogeographies (c) for Triassic (T), Jurassic (J) and Cretaceous (K). Silhouette colours symbolize body mass for each of the taxa represented; information on dietary habits are plotted after Barrett and Zanno & Makovicky; numbers represent clades discussed through this study: 1, Ornithischia; 2, Thyreophora; 3, Ornithopoda; 4, Hadrosauroidea; 5, Marginocephalia; 6, Ceratopsia; 7, Saurischia; 8, Sauropodomorpha; 9, Sauropoda; 10, Theropoda; 11, Ceratosauria; 12, Tetanurae; 13, Coelurosauria; 14, Maniraptoriformes; 15, Maniraptora; 16, Deinonychosauria; 17, Avialae; 18, Ornithothoraces. Palaeogeographies modified from original plots via R package ‘mapast’ using plate models by Scotese.

Diagram illustrating the "Temporal paradox" in paleontology. First given it's nickname by Alan Feduccia, the paradox is made up by the fact that almost all feathered dinosaurs are dated to have lived millions of years after Archaeopteryx, the oldest bird (late Jurassic, believed to have existed about 150 million years ago). Only a few of the feathered dinosaurs/birdlike dinosaurs are given an older date than Archaeopteryx.

Montage of eight different representatives of coelurosaurian dinosaurs: (Clockwise from upper left) GIN 100/42 which may represent Citipati or a different oviraptorosaur, Sinosauropteryx prima, Nothronychus mckinleyi, Tyrannosaurus rex, Bambiraptor feinbergi, Passer domesticus, Struthiomimus altus, Microraptor gui. This is a collection of eight different works already found in Wikimedia Commons (see source field below).

Simplified cladogram of Dinosauria with the distribution of feathers according to the fossil record. Despite its more ancient origin, it was only in maniraptoriformes that modern-type feathers (pennaceous feathers) have arisen (Based in Xu & Guo, 2009; Clarke, 2013; Godefroit et al., 2013; Han et al., 2014; Koshchowitz et al., 2014).

Ареал карнозавров

Reconstruction of the theropod dinosaur Tachiraptor admirabilis. Artwork created with graphite and colored pencils. Author: Edwin Chávez "Disfrasaurio".

Ceratosaurus nasicornis reconstruction. Matches proportions shown in Gregory S. Paul (The Princeton Field Guide to Dinosaurs, 2010, p. 84)

Sketch-drawing of the vertebrate faunal assemblage of the Lisowice site (modified from Niedźwiedzki)10. (a) Large, theropod-like predatory archosaur (Smok wawelski); (b) large temnospondyl amphibian (Cyclotosaurus sp.); (c) small predatory dinosaurs (Neotheropoda indet.); (d) temnospondyl amphibian (Gerrothorax sp.); (e) small basal crocodylomorph (Crocodylomorpha indet.); (f) small diapsid (Choristodere-like animal); (g) hybodont sharks (Polyacrodus and Hybodus); (h) coelacanth fish; (i) dipnoan fish (Ptychoceratodus sp.); (j) actinopterygian fish; (k) gigantic dicynodont; (l) dinosauriforms or early dinosaurs (Dinosauriformes indet. or Dinosauria indet.); (m) small lepidosauromorphs (Sphenodontia indet.); (n) pterosaurs (Pterosauria indet.); (o) early mammaliaform (Hallautherium sp.).

Diagram made to illustrate the debate over the hands of theropod dinosaurs and their alleged descendants, the birds ( Aves ). In the last years, the difference between the hands of theropods and birds have been an important thing when it comes question the theory that birds evolved from dinosaurs, even though some scientists may could refute it. In the diagram, a Neotheropoda ( 1 ), basal tetanurae ( 2 ), a coelurosaurian ( 3 ), the bird (?)Archaeopteryx ( 4 ) and modern bird ( 5 ). In 1997, birdexpert Alan Feduccia at University of North Carolina discovered that birds develop hands with the digits II, III and IV ( see The Cincinnati Enquirer, 25 - 10 - 1997 ). This is in contrast with the hands of tetanurae, which seems to have the digits I, II and III. This make it almost impossible for dinosaurs and birds to be closely related, according to Feduccia. Since the discovery by Feduccia, scientific research have came up with a possible explanation to the mystery of the dinosaur - bird hand difference, called The frame shift hypothesis ( see http://scienceblogs.com/tetrapodzoology/2009/06/limusaurus_is_awesome.php ). This hypothesis is based on a discovery which shows that although bird embryos develop the fingers II, III and IV, the genes which is coding for the external appearance of the digits seems to be from the fingers I, II and III. Based on this, scientists belive this: when neotheropods evolved into tetanurae, the losed digit I ( not digit IV, as earlier suggested ). During this process, the genes which coded for how the digits should looks like ( the number of phalanges, for example ) became refurnished ( see the color spots in th upper section in the diagram to understan ). The discovery of Limusaurus has been said to support this theory ( see https://www.livescience.com/animals/090617-dinosaur-hands.html ). One thing is that may can be used to refute that Limusaurus should support the Frame shift hypothesis is that Limusaurus was a ceratosaurian, and is dated to be much younger than the oldest tetanurae's. Also, some tetanure´s may had 4 digits' like the Archaeornithomimus ( see number 4 in the diagram ) ( see also http://dml.cmnh.org/1998Oct/msg00443.html and the Allosaurus hand in the image here: http://upload.wikimedia.org/wikipedia/commons/6/69/Allosaurus-mounted.jpg ). If this is the digits I, II, III and IV. If so, it shows that tetanurae had the digits I, II and III, and not II, II and IV, like birds. For more in this debate, see text section to my picture Raptor-Archaeopteryx-bird hands differens.JPG at http://commons.wikimedia.org/wiki/File:Raptor-Archaeopteryx-bird_hands_differens.JPG.

Cladogram of amniotes. Based on File:Cladogram Amniota A.jpg

Cladogram of the former "Reptilia" (red)

Dinosaur art by me!