Cretaceous

Geological interval

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Azhdarchoid pterosaur Kariridraco dianae gen. et sp. nov. holotype (MPSC R 1056) from Lower Cretaceous of Northeastern Brazil. Skull in right lateral view. Photograph (A1), interpretative drawing (A2). Dark grey areas indicate remains of the carbonate matrix still attachaed to the skull.
Taxa Kariridraco

Azhdarchoid pterosaur Kariridraco dianae gen. et sp. nov. holotype (MPSC R 1056) from Lower Cretaceous of Northeastern Brazil. Skull in right lateral view. Photograph (A1), interpretative drawing (A2). Dark grey areas indicate remains of the carbonate matrix still attachaed to the skull.

drawing Brazil Cretaceous holotype +3
Abelisaurid theropod Kryptops palaios gen. et sp. nov. MNN GAD1−1 from the Lower Cretaceous Elrhaz Formation of Niger. Left maxilla in lateral

view; stereopair (A) and line drawing (B). Cross−hatching indicates broken bone; dashed lines indicate estimated edges.
Taxa Kryptops

Abelisaurid theropod Kryptops palaios gen. et sp. nov. MNN GAD1−1 from the Lower Cretaceous Elrhaz Formation of Niger. Left maxilla in lateral view; stereopair (A) and line drawing (B). Cross−hatching indicates broken bone; dashed lines indicate estimated edges.

bone drawing Niger Cretaceous +3
Hypothetical reconstruction of the Late Cretaceous abelisaurid Elemgasem nubilus.
Taxa Elemgasem

Hypothetical reconstruction of the Late Cretaceous abelisaurid Elemgasem nubilus.

Cretaceous Late Cretaceous Abelisauridae Elemgasem
Riojavenatrix: "La Rioja huntress" Early Cretaceous, Europe
Taxa Riojavenatrix

Riojavenatrix: "La Rioja huntress" Early Cretaceous, Europe

Cretaceous Early Cretaceous Riojavenatrix
Left maxilla (A) and left dentary (B) of carcharodontosaurid theropod Kelmayisaurus petrolicus Dong, 1973 (IVPP V 4022, China, Lianmugin For−
mation, Lower Cretaceous). Photographs in lateral (A1, B1), medial (A2, B2), ventral (A3), and dorsal (B3) views. Scale bars 5 cm. Designation “d” refers to

dentary tooth, designation “m” refers to maxillary tooth position. Dorsal view of dentary shows tooth row only.
Taxa Kelmayisaurus

Left maxilla (A) and left dentary (B) of carcharodontosaurid theropod Kelmayisaurus petrolicus Dong, 1973 (IVPP V 4022, China, Lianmugin For− mation, Lower Cretaceous). Photographs in lateral (A1, B1), medial (A2, B2), ventral (A3), and dorsal (B3) views. Scale bars 5 cm. Designation “d” refers to dentary tooth, designation “m” refers to maxillary tooth position. Dorsal view of dentary shows tooth row only.

scale tooth China Cretaceous +2
Closeup of the camellate pneumaticity inside the vertebrae of Vectiraptor greeni, from the Early Cretaceous of the Isle of Wight, UK. Photo by Nick Longrich.
Taxa Vectiraptor

Closeup of the camellate pneumaticity inside the vertebrae of Vectiraptor greeni, from the Early Cretaceous of the Isle of Wight, UK. Photo by Nick Longrich.

United Kingdom Cretaceous Early Cretaceous Vectiraptor
Diagram featuring the holotype (BNMNH-PV030) skull of the troodontid Papiliovenator neimengguensis, from the Bayan Mandahu Formation. Abbreviations: a, angular; d, dentary; f, frontal; j, jugal; l, lacrimal; m, maxilla; n, nasal; p, parietal; q, quadrate;  s, surangular;  sp, splenial.[1]
References

↑  (2022). "A new troodontid from the Upper Cretaceous Gobi Basin of inner Mongolia, China". Cretaceous Research 130: Article 105052. DOI:10.1016/j.cretres.2021.105052.
Taxa Papiliovenator

Diagram featuring the holotype (BNMNH-PV030) skull of the troodontid Papiliovenator neimengguensis, from the Bayan Mandahu Formation. Abbreviations: a, angular; d, dentary; f, frontal; j, jugal; l, lacrimal; m, maxilla; n, nasal; p, parietal; q, quadrate; s, surangular; sp, splenial.[1] References ↑ (2022). "A new troodontid from the Upper Cretaceous Gobi Basin of inner Mongolia, China". Cretaceous Research 130: Article 105052. DOI:10.1016/j.cretres.2021.105052.

China Mongolia Cretaceous holotype +4
Skull of troodontid theropod Xixiasaurus henanensis gen. et sp. nov. (HGM 41HIII−0201; holotype), lower–middle Majiacun Formation (Upper Cretaceous: Coniacian–Campanian), Henan Province, China; in dorsal (A); lateral (B), and ventral (C) views.

Note that this version of the image does not include the interpretative lines of the version in the paper.
Taxa Xixiasaurus

Skull of troodontid theropod Xixiasaurus henanensis gen. et sp. nov. (HGM 41HIII−0201; holotype), lower–middle Majiacun Formation (Upper Cretaceous: Coniacian–Campanian), Henan Province, China; in dorsal (A); lateral (B), and ventral (C) views. Note that this version of the image does not include the interpretative lines of the version in the paper.

China Campanian Coniacian Cretaceous +5
Life reconstruction of the Late Cretaceous south american paravian, Overoraptor chimentoi
Taxa Overoraptor

Life reconstruction of the Late Cretaceous south american paravian, Overoraptor chimentoi

Cretaceous Late Cretaceous Overoraptor
Forearm and hand of Machairasaurus leptorhynchos Longrich et al. 2010 (Dinosauria: Theropoda: Oviraptoridae: Ingeniinae) from the Late Cretaceous Djadokhta Formation beds of Bayan Mandahu, Inner Mongolia, China. Digital photography and editing by Nick Longrich
Taxa Machairasaurus

Forearm and hand of Machairasaurus leptorhynchos Longrich et al. 2010 (Dinosauria: Theropoda: Oviraptoridae: Ingeniinae) from the Late Cretaceous Djadokhta Formation beds of Bayan Mandahu, Inner Mongolia, China. Digital photography and editing by Nick Longrich

China Mongolia Djadokhta Cretaceous +8
Alvarezsauridae gen. et sp. indet., posterior caudal vertebrae. Dzharakuduk, Uzbekistan; Bissekty Formation, Upper Cretaceous (Turonian). a-e, ZIN PH 2441/16, two vertebrae preserved in articulation, in anterior (a), dorsal (b), posterior (c), lateral (d), and ventral (e) views. f-j, ZIN PH 2442/16, vertebra missing ventral part of the centrum, in dorsal (f), lateral (g), ventral (h), anterior (i), and posterior (j).
Taxa Dzharaonyx

Alvarezsauridae gen. et sp. indet., posterior caudal vertebrae. Dzharakuduk, Uzbekistan; Bissekty Formation, Upper Cretaceous (Turonian). a-e, ZIN PH 2441/16, two vertebrae preserved in articulation, in anterior (a), dorsal (b), posterior (c), lateral (d), and ventral (e) views. f-j, ZIN PH 2442/16, vertebra missing ventral part of the centrum, in dorsal (f), lateral (g), ventral (h), anterior (i), and posterior (j).

vertebra Uzbekistan Bissekty Cretaceous +4
Ceratonykus oculatus, Late Cretaceous of Mongolia. Digital.
Taxa Ceratonykus

Ceratonykus oculatus, Late Cretaceous of Mongolia. Digital.

Mongolia Cretaceous Late Cretaceous Ceratonykus +1
Speculative reconstruction of Aoniraptor'’ from the Late Cretaceous of Argentina, here restored as a megaraptoran as recovered by most studies
Taxa Aoniraptor

Speculative reconstruction of Aoniraptor'’ from the Late Cretaceous of Argentina, here restored as a megaraptoran as recovered by most studies

Argentina Cretaceous Late Cretaceous Aoniraptor +1
Skull diagram featuring the holotype skull (ROM 1790) of the ornithomimid Rativates evadens, from the Late Cretaceous period. Abbreviations: d, dentary; f, frontal; l, lacrimal; m, maxilla; n, nasal; pm, premaxila; ps, paraphenoid.
Taxa Rativates

Skull diagram featuring the holotype skull (ROM 1790) of the ornithomimid Rativates evadens, from the Late Cretaceous period. Abbreviations: d, dentary; f, frontal; l, lacrimal; m, maxilla; n, nasal; pm, premaxila; ps, paraphenoid.

Cretaceous Late Cretaceous holotype Ornithomimidae +2
Ceratopsipes goldenensis Lockley & Hunt, 1995 - ceratopsian dinosaur trackway in the Cretaceous of Colorado, USA.
Ceratopsians are the "horned dinosaurs".  They were large, quadrupedal, herbivorous dinosaurs having a beaked skull and a frill - an extension of bone behind the skull that partially covered the neck.  Ceratopsian dinosaurs are known from the Jurassic and Cretaceous.  The last members of the group died out at the Cretaceous-Tertiary boundary, 65 million years ago.
Seen here is a ceratopsian dinosaur trackway.  Ceratopsian footprints are very rare.  The best examples are in Colorado's Laramie Formation, a nonmarine, coastal plain to deltaic succession of mostly mixed siliciclastic sedimentary rocks - sandstones, siltstones, claystones, and coals.  The footprints here consist of convex bulges on the basal surfaces of structurally-tilted sandstone beds.
The ichnospecies Ceratopsipes goldenensis was named based on footprints at this locality.  The trackmaker was very likely Triceratops, the # 1 most famous ceratopsian dinosaur.


From on-site signage:
Triceratops Tracks
Several tracks of Triceratops, or a closely related horned dinosaur were first discovered in this area and named Ceratopsipes goldenensis (meaning tracks of a horned dinosaur from Golden).  By happy coincidence, the first Triceratops known to science also comes from the greater Denver area near 13th Avenue and Federal Blvd.  Growing to a length of almost 25 feet, Triceratops and other members of the horned dinosaur family were herbivorous animals that roamed in herds.  Their trademark horns and frills that covered their necks probably served for defense.  A replica Triceratops skull is on display in the clubhouse at the Fossil Trace Golf Club.
These tracks (identified by small signs) help us understand that Triceratops walked with a narrow gait and erect front limbs  than with a wide, sprawling gait as previously depicted.  In 1887, the first Triceratops bones known to science were also discovered locally at a site near 13th Avenue and Federal Boulevard in Denver.
Nearby one may see many other traces of life such as burrows, probably made by invertebrates and impressions of plant debris.  Note that you are looking at all tracks and traces from the underside - in negative aspect - what one might call a worm’s eye view.  All of these trace fossils indicate that the Golden area once had a warm, temperate to subtropical climate.


Classification: Animalia, Chordata, Vertebrata, Reptilia, Archosauria, Dinosauria, Ornithischia, Marginocephalia, Ceratopsia, Ceratopsidae
Stratigraphy: Laramie Formation, Maastrichtian Stage, upper Upper Cretaceous
Locality: outcrop along Triceratops Trail, Parfet Prehistoric Preserve, southern side of the town of Golden, Colorado, USA (~vicinity of 39° 44' 35.24" North latitude, 105° 13’ 09.69" West longitude)


Some info. from:

Lockley & Hunt (1995) - Ceratopsid tracks and associated ichnofauna from the Laramie Formation (Upper Cretaceous: Maastrichtian) of Colorado.  Journal of Vertebrate Paleontology 15: 592-614.
Taxa Ceratopsipes

Ceratopsipes goldenensis Lockley & Hunt, 1995 - ceratopsian dinosaur trackway in the Cretaceous of Colorado, USA. Ceratopsians are the "horned dinosaurs". They were large, quadrupedal, herbivorous dinosaurs having a beaked skull and a frill - an extension of bone behind the skull that partially covered the neck. Ceratopsian dinosaurs are known from the Jurassic and Cretaceous. The last members of the group died out at the Cretaceous-Tertiary boundary, 65 million years ago. Seen here is a ceratopsian dinosaur trackway. Ceratopsian footprints are very rare. The best examples are in Colorado's Laramie Formation, a nonmarine, coastal plain to deltaic succession of mostly mixed siliciclastic sedimentary rocks - sandstones, siltstones, claystones, and coals. The footprints here consist of convex bulges on the basal surfaces of structurally-tilted sandstone beds. The ichnospecies Ceratopsipes goldenensis was named based on footprints at this locality. The trackmaker was very likely Triceratops, the # 1 most famous ceratopsian dinosaur. From on-site signage: Triceratops Tracks Several tracks of Triceratops, or a closely related horned dinosaur were first discovered in this area and named Ceratopsipes goldenensis (meaning tracks of a horned dinosaur from Golden). By happy coincidence, the first Triceratops known to science also comes from the greater Denver area near 13th Avenue and Federal Blvd. Growing to a length of almost 25 feet, Triceratops and other members of the horned dinosaur family were herbivorous animals that roamed in herds. Their trademark horns and frills that covered their necks probably served for defense. A replica Triceratops skull is on display in the clubhouse at the Fossil Trace Golf Club. These tracks (identified by small signs) help us understand that Triceratops walked with a narrow gait and erect front limbs than with a wide, sprawling gait as previously depicted. In 1887, the first Triceratops bones known to science were also discovered locally at a site near 13th Avenue and Federal Boulevard in Denver. Nearby one may see many other traces of life such as burrows, probably made by invertebrates and impressions of plant debris. Note that you are looking at all tracks and traces from the underside - in negative aspect - what one might call a worm’s eye view. All of these trace fossils indicate that the Golden area once had a warm, temperate to subtropical climate. Classification: Animalia, Chordata, Vertebrata, Reptilia, Archosauria, Dinosauria, Ornithischia, Marginocephalia, Ceratopsia, Ceratopsidae Stratigraphy: Laramie Formation, Maastrichtian Stage, upper Upper Cretaceous Locality: outcrop along Triceratops Trail, Parfet Prehistoric Preserve, southern side of the town of Golden, Colorado, USA (~vicinity of 39° 44' 35.24" North latitude, 105° 13’ 09.69" West longitude) Some info. from: Lockley & Hunt (1995) - Ceratopsid tracks and associated ichnofauna from the Laramie Formation (Upper Cretaceous: Maastrichtian) of Colorado. Journal of Vertebrate Paleontology 15: 592-614.

bone United States Cretaceous Jurassic +6
Ornithischian dinosaur Stenopelix valdensis Meyer, 1857, holotype (GZG 741/2, formerly GPI Gö 741−2), from the Obernkirchen Sandstone (Early
Cretaceous: Berriasian), near Bückeburg, Niedersachsen, Germany. A. Large sandstone slab. B. Interpretative outline drawing of large latex cast (prepared
from large sandstone slab), showing majority of postcranial skeleton in dorsal view. C. Small sandstone slab. D. Interpretative outline drawing of small la−
tex cast (prepared from small sandstone slab), showing sacrum and caudals, pelvic region and partial hindlimbs in ventral view. For clarity elements in and
around the sacral region have not been labelled—these areas are shown in greater detail in Fig. 3. Roman numerals II–IV correspond to respective digits.

Abbreviations: mt, metatarsals; f, femur; ?, unidentified element.
Taxa Stenopelix

Ornithischian dinosaur Stenopelix valdensis Meyer, 1857, holotype (GZG 741/2, formerly GPI Gö 741−2), from the Obernkirchen Sandstone (Early Cretaceous: Berriasian), near Bückeburg, Niedersachsen, Germany. A. Large sandstone slab. B. Interpretative outline drawing of large latex cast (prepared from large sandstone slab), showing majority of postcranial skeleton in dorsal view. C. Small sandstone slab. D. Interpretative outline drawing of small la− tex cast (prepared from small sandstone slab), showing sacrum and caudals, pelvic region and partial hindlimbs in ventral view. For clarity elements in and around the sacral region have not been labelled—these areas are shown in greater detail in Fig. 3. Roman numerals II–IV correspond to respective digits. Abbreviations: mt, metatarsals; f, femur; ?, unidentified element.

drawing Germany Berriasian Cretaceous +8
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bone Japan Cretaceous fossil Canardia Dinosauria Iguanodontia
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bone jaw Cretaceous fossil Mosasaurus new species skull
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bone Mexico Campanian Cretaceous Late Cretaceous fossil Dinevenator Dinosauria Troodontidae new species skull
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predator prey Australia Toolebuc Cretaceous Early Cretaceous fossil Ichthyosauria Pterosauria formation
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Myanmar Cretaceous new species other reptiles
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